Taxonomy of Lucanidae

Subfamily Aesalinae :Tribe Aesalini ::Genus Aesalus (Fabricius, 1801) :::*Aesalus asiaticus Lewis, 1883 :::*Aesalus himalayicus Kurosawa, 1985 :::*Aesalus imanishii Inahara & Ratti, 1991 :::*Aesalus meridionalis Bartolozzi, 1989 :::*Aesalus neotropicalis Bates, 1886 :::*Aesalus saburoi :::*Aesalus sawaii Fujita & Ichikawa, 1985 :::*Aesalus scarabaeoides Panzer, 1793 :::*Aesalus sichuanensis :::*Aesalus smithi Bates, 1889 :::*Aesalus trogoides Albers, 1883 :::*Aesalus ulanowskii Ganglbauer, 1886 ::Genus Cretaesalus (fossil) ::Genus Echinoaesalus Zelenka, 1993 :::*Echinoaesalus barriesi (Zelenka, 1993) :::*Echinoaesalus hidakai (Araya, 1993) :::*Echinoaesalus jaechi (Zelenka,1993) :::*Echinoaesalus matsuii (Araya, 1993) :::*Echinoaesalus sabahensis (Zelenka, 1994) :::*Echinoaesalus schuhi (Zelenka, 1994) :::*Echinoaesalus timidus (Krikken, 1975) :::*Echinoaesalus yongi (Araya, 1993) ::Genus Lucanobium Howden & Lawrence, 1974 :::*Lucanobium squamosum Howden & Lawrence, 1974 :Tribe Ceratognathini ::Genus Ceratognathus Westwood, 1838 :::*Ceratognathus abdominalis Parry, 1870 :::*Ceratognathus bitumulatus Carter, 1925 :::*Ceratognathus flabellatus Boileau, 1905 :::*Ceratognathus frenchi Blackburn, 1897 :::*Ceratognathus froggatti Blackburn, 1894 :::*Ceratognathus gilesi Blackburn, 1895 :::*Ceratognathus macrognathus Boileau, 1905 :::*Ceratognathus mentiferus Westwood, 1863 :::*Ceratognathus minutus Lea, 1929 :::*Ceratognathus niger Westwood, 1838 :::*Ceratognathus ocularis Carter, 1925 :::*Ceratognathus rufipennis Westwood, 1872 :::*Ceratognathus tasmanus Benesh, 1943 :::*Ceratognathus westwoodi Thomson, 1862 ::Genus Hilophyllus Paulsen & Mondaca, 2006 :::*Hilophyllus argentinensis (Martinez, 1981) :::*Hilophyllus martinezi Paulsen & Mondaca, 2006 :::*Hilophyllus penai (Martinez, 1976) ::Genus Holloceratognathus Nikolajev, 1998 :::*Holloceratognathus cylindricus (Broun, 1895) :::*Holloceratognathus helotoides (Thomson, 1862) :::*Holloceratognathus passaliformis (Holloway, 1962) ::Genus Mitophyllus Parry, 1843 :::*Mitophyllus alboguttatus (Bates, 1867) :::*Mitophyllus dispar (Sharp, 1882) :::*Mitophyllus foveolatus (Broun, 1880) :::*Mitophyllus gibbosus (Broun, 1884) :::*Mitophyllus insignis Broun, 1921 :::*Mitophyllus irroratus Parry, 1842 :::*Mitophyllus macrocerus (Broun, 1886) :::*Mitophyllus parrianus Westwood, 1863 :::*Mitophyllus reflexus Broun, 1908 :Tribe Nicagini ::Genus Nicagus LeConte, 1861 :::*Nicagus obscurus (LeConte, 1848) :::*Nicagus occultus Paulsen & Smith, 2005 :::*Nicagus japonicus Nagel, 1928 ==Subfamily Lampriminae==

Subfamily Lampriminae ::Genus Dendroblax White, 1846 :::*Dendroblax earlianus White, 1846 ::Genus Homolamprima :::*Homolamprima crenulata (MacLeay, 1885) ::Genus Lamprima :::*Lamprima adolphinae (Gestro, 1875) :::*Lamprima aenea Fabricius, 1792 :::*Lamprima aurata Latreille, 1817 :::*Lamprima imberbis Barber, 1926 :::*Lamprima insularis Macleay, 1885 ::Genus Phalacrognathus :::*Phalacrognathus muelleri MacLeay, 1885 ::Genus Streptocerus Dejean, 1833 :::*Streptocerus speciosus Fairmaire, 1850 ==Subfamily Lucaninae==

Subfamily Lucaninae Latreille, 1804 :Tribe Aegini ::Genus Aegus MacLeay, 1819 :Tribe Allotopini ::Genus Allotopus :::*Allotopus moellenkampi (Fruhstorfer, 1894) :::*Allotopus rosenbergi (Vollenhoven, 1872) ::Genus Mesotopus :Tribe Chiasognathini ::Genus Casignetus ::Genus Chiasognathus Stephens, 1831 :::*Chiasognathus beneshi Lacroix, 1978 :::*Chiasognathus grantii Stephens, 1831 :::*Chiasognathus jousselini Reiche, 1850 :::*Chiasognathus latreillei Solier, 1851 :::*Chiasognathus mniszechii Kiesche 1919 ::Genus Protognathinus ::Genus Sphaenognathus :Tribe Cladognathini ::Genus Aphanognathus :::*Aphanognathus laterotarsoides (Houlbert, 1915) ::Genus Capreolucanus :::*Capreolucanus sicardi Didier, 1928 ::Genus Cladophyllus ::Genus Eligmodontus ::Genus Gonometopus ::Genus Macrodorcas Motschulsky, 1862 :::*Macrodorcas adachii :::*Macrodorcas amamiana :::*Macrodorcas barbata :::*Macrodorcas bisignata :::*Macrodorcas bomansi :::*Macrodorcas davidi :::*Macrodorcas dierli :::*Macrodorcas elsiledis :::*Macrodorcas fulvonotata :::*Macrodorcas humilis :::*Macrodorcas intricata :::*Macrodorcas kobayashii :::*Macrodorcas koyamai :::*Macrodorcas melliana :::*Macrodorcas mochizukii :::*Macrodorcas moellenkampi :::*Macrodorcas montivaga :::*Macrodorcas pieli :::*Macrodorcas pseudaxis :::*Macrodorcas recta Motschulsky, 1858 :::*Macrodorcas ruficrus :::*Macrodorcas songiana :::*Macrodorcas striatipennis Motschulsky, 1862 :::*Macrodorcas ursulae :::*Macrodorcas vernicata :::*Macrodorcas yaeyamaensis ::Genus Palaeognathus ::Genus Prismognathus Motschulsky, 1860 :::*Prismognathus alessandrae :::*Prismognathus angularis Waterhouse, 1874 :::*Prismognathus branczicki :::*Prismognathus cheni :::*Prismognathus dauricus Motschulsky, 1860 :::*Prismognathus davidis :::*Prismognathus delislei :::*Prismognathus formosanus :::*Prismognathus klapperichi :::*Prismognathus lucidus :::*Prismognathus morimotoi :::*Prismognathus nigerrimus :::*Prismognathus nigricolor :::*Prismognathus piluensis :::*Prismognathus platycephalus :::*Prismognathus ruficephalus :::*Prismognathus sinensis :::*Prismognathus subnitens :::*Prismognathus tokui ::Genus Prosopocoilus Westwood, 1845 ::Genus Pseudorhaetus ::Genus Rhaetulus ::Genus Rhaetus :::*Rhaetus westwoodi (Parry, 1862) ::Genus Tetrarthrius ::Genus Weinreichius :::*Weinreichius perroti Lacroix, 1978 :Tribe Colophonini ::Genus Colophon :Tribe Cyclommatini ::Genus Cyclommatus Parry, 1863 :Tribe Dendeziini ::Genus Dendezia ::Genus Oonotus Parry, 1864 ::Genus Xiphodontus :Tribe Dorcini ::Genus Bartolozziolucanus ::Genus Cantharolethrus :::*Cantharolethrus luxeri ::Genus Cyclommatus :::*Cyclommatus scutellaris ::Genus Dorcasoides ::Genus Dorcus MacLeay, 1819 :::*Dorcus akahorii :::*Dorcus alcides :::*Dorcus alexisi :::*Dorcus amamianus :::*Dorcus antaeus :::*Dorcus anteus :::*Dorcus arfakianus :::*Dorcus binodulosus :::*Dorcus brevis :::*Dorcus bucephalus :::*Dorcus carinulatus :::*Dorcus consentaneus :::*Dorcus curvidens :::*Dorcus cylindricus :::*Dorcus davidis :::*Dorcus emikoae :::*Dorcus eurycephalus :::*Dorcus formosanus :::*Dorcus gracilicornis :::*Dorcus garndis :::*Dorcus hopei :::*Dorcus hyperion :::*Dorcus japonicus :::*Dorcus kyanrauensis :::*Dorcus metacostatus :::*Dorcus miwai :::*Dorcus musimon :::*Dorcus parallelus :::*Dorcus parallelipipedus - lesser stag beetle. Linnaeus, 1758 :::*Dorcus parryi :::*Dorcus parvulus :::*Dorcus peyronis :::*Dorcus prochazkai :::*Dorcus ratiocinativus :::*Dorcus rectus :::*Dorcus reichei :::*Dorcus rudis :::*Dorcus sawaii :::*Dorcus schenklingi :::*Dorcus sewertzowi :::*Dorcus suturalis :::*Dorcus taiwanicus :::*Dorcus tanakai :::*Dorcus taurus :::*Dorcus tenuecostatus :::*Dorcus titanus :::*Dorcus tormosanus :::*Dorcus ursulus :::*Dorcus vavrai :::*Dorcus velutinus :::*Dorcus vicinus :::*Dorcus yamadai ::Genus Homoderus ::Genus Leptinopterus ::Genus Prosopocoilus :::*Prosopocoilus astacoides :::*Prosopocoilus biplagiatus :::*Prosopocoilus bison :::*Prosopocoilus giraffa :::*Prosopocoilus inclinatus :::*Prosopocoilus savagei ::Genus Rhaetulus :::*Rhaetulus crenatus ::Genus Serrognathus ::Genus Pycnosiphorus ::Genus Sclerostomus :Tribe Figulini ::Genus Cardanus ::Genus Dinonigidius ::Genus Figulus ::Genus Ganelius ::Genus Nigidionus ::Genus Nigidius ::Genus Novonigidius ::Genus Penichrolucanus :Tribe Lissapterini ::Genus Bomansius ::Genus Dorculus ::Genus Geodorcus ::Genus Hoplogonus ::Genus Lissapterus ::Genus Lissotes ::Genus Paralissotes ::Genus Pseudodorcus :Tribe Lucanini ::Genus Hexarthrius Hope, 1842 :::*Hexarthrius aduncusJordan & Rothschild, 1894 :::*Hexarthrius andreasi Schenk, 2003 :::*Hexarthrius bowringii Parry, 1862 :::*Hexarthrius buquettii (Hope, 1843) :::*Hexarthrius davisoniWaterhouse, 1888 :::*Hexarthrius forsteri (Hope, 1840) :::*Hexarthrius howdeni De Lisle, 1972 :::*Hexarthrius kirchneri Schenk, 2003 :::*Hexarthrius mandibularis Deyrolle, 1881 :::*Hexarthrius melchioritis Séguy, 1954 :::*Hexarthrius mniszechi(Thomson, 1857) :::*Hexarthrius nigritus Lacroix, 1990 :::*Hexarthrius parryi Hope, 1842 :::*Hexarthrius rhinoceros(Olivier, 1789) :::*Hexarthrius vitalisi Didier, 1925 ::Genus Lucanus :::*Lucanus capreolus :::*Lucanus cervus :::*Lucanus elaphus :::*Lucanus formosanus :::*Lucanus maculifemoratus :::*Lucanus mazama :::*Lucanus placidus :::*Lucanus swinhoei :::*Lucanus tetraodon ::Genus Pseudolucanus :::*Pseudolucanus barbarossa (Fabricius, 1801) :Tribe Odontolabini ::Genus Calcodes ::Genus Neolucanus :::*Neolucanus castanopterus ::Genus Odontolabis :::*Odontolabis castelnaudi :::*Odontolabis cuvera :::*Odontolabis femoralis :::*Odontolabis siva :Tribe Platycerini ::Genus Platyceroides Benesh, 1946 ::Genus Platyceropsis Benesh, 1946 :::*Platyceropsis keeni (Casey, 1895) ::Genus Platycerus Geoffroy, 1762 :Tribe Ryssonotini ::Genus Cacostomus :::*Cacostomus floralis (Lea, 1914) :::*Cacostomus squamosus Newman, 1840 :::*Cacostomus subvittata (Moore, 1994) ::Genus Ryssonotus ==Subfamily Syndesinae==

Subfamily Syndesinae :Tribe Ceruchini ::Genus Ceruchus MacLeay, 1819 :::*Ceruchus piceus (Weber, 1801) :::*Ceruchus punctatus LeConte, 1869 :::*Ceruchus striatus LeConte, 1859 :::*Ceruchus yangi Huang, Imura & Chen, 2011 :Tribe Sinodendronini ::Genus Sinodendron Hellwig in Schneider, 1792 :::*Sinodendron cylindricum (Linnaeus, 1758) :::*Sinodendron rugosum Mannerheim, 1843 :Tribe Syndesini ::Genus Psilodon Perty, 1830 :::*Psilodon aequinoctiale (Buquet, 1840) :::*Psilodon gilberti Boucher, 1993 :::*Psilodon schuberti Perty, 1830 :::*Psilodon seguyi (Didier, 1929) :::*Psilodon xerophilicum Martinez & Reyes-Castillo, 1985 ::Genus Syndesus MacLeay, 1819 :::*†Syndesus ambericus Woodruff, 2009 (Miocene?; Dominican amber) :::*Syndesus cancellatus Montrouzier, 1860 :::*Syndesus cornutus (Fabricius, 1801) :::*Syndesus macleayi Boileau, 1905 == References ==