The subspecies
Ensatina e. eschscholtzii, the Monterey ensatina, can be found in
Santa Cruz and
Monterey Counties and into the
California coastal mountains. With a head-to-tail length of just between ,
E. e. eschecholtzi can be identified primarily by its tail, which is narrower at the base; it is the only subspecies that has such a tail structure, as well as five toes on the hind limbs. Males often have longer tails than the females, and many of these salamanders have lighter-colored limbs, compared to the rest of the body. The adult females lay eggs underground, often in sets of threes, which hatch directly into fully-formed salamanders, skipping the usual aquatic juvenile phase. The subspecies
Ensatina eschscholtzii klauberi, or the large-blotched ensatina, can be found along the mountain ranges of
Southern California, and south into a small region of the Sierra Juarez in northern
Baja California.
E. e. klauberi is similar in size to
E. e. eschscholtzii; it is mid-sized, with adults growIng a total length of . Females tend to have shorter, wider bodies compared to the males. However, this subspecies differs from
E. e. eschscholtzii in its coloration—nearly black, with blotches of orange, tail, and dark eyes.
As a ring species Ensatina eschscholtzii has been described as a
ring species in the mountains surrounding the
Californian Central Valley. The complex population forms a
horseshoe shape around the mountains, and although
interbreeding can happen between each of the 19 populations around said horseshoe, the
Ensatina eschscholtzii subspecies on the western end of the horseshoe cannot interbreed with the
Ensatina klauberi on the eastern end. As such, it is thought to be an example of
incipient speciation, providing an illustration of "nearly all stages in a speciation process" (Dobzhansky, 1958).
Richard Highton,
zoologist, argued that
Ensatina is a genus of multiple species and not a continuum of one (meaning, by traditional definitions, it is not a ring species). ==Distribution and habitat==