The genus
Haliaeetus was introduced in 1809 by French naturalist
Marie Jules César Savigny in his chapter on birds in the ''
Description de l'Égypte. In 2005, Haliaeetus was found to be paraphyletic after molecular study was performed; that genus was found to subsume Icthyophaga with the species within it diverging into temperate and tropical groups. Subsequently, the two species of Icthyophaga
were accordingly moved within Haliaeetus'', within the tropical group. However, an academic paper published in 2024 based on a densely sampled
molecular phylogenetic study of the Accipitridae by Therese Catanach and collaborators found
Icthyophaga to be distinct enough to be their own genus from
Haliaeetus. This resulted in
Frank Gill,
Pamela C. Rasmussen and David Donsker on behalf of the
International Ornithological Committee (IOC) resurrecting
Icthyophaga, and also moving the more tropical members of
Haliaeetus into a now-expanded
Icthyophaga.
Species Evolution Haliaeetus is possibly one of the oldest genera of living birds, remaining
extant until today. A
distal left
tarsometatarsus (DPC 1652) recovered from early
Oligocene deposits of Fayyum,
Egypt (
Jebel Qatrani Formation, about 33 million years ago (Mya)) is similar in general pattern and some details to that of a modern sea eagle. The genus was present in the middle
Miocene (12–16 Mya) with certainty. The point of origin of the sea- and fishing eagles is probably in the general area of the
Bay of Bengal. During the
Eocene/
Oligocene, as the
Indian subcontinent slowly collided with
Eurasia, the region was a vast expanse of fairly shallow ocean; the initial sea eagle
divergence seems to have resulted in the four
tropical (and
Southern Hemisphere subtropical) species living around the
Indian Ocean today. The
Central Asian Pallas's sea eagle's relationships to the other
taxa is more obscure; it seems closer to the three
Holarctic species which evolved later and may be an early offshoot of this northward expansion; it does not have the hefty yellow bill of the northern forms, retaining a smaller, darker beak like the tropical species. A prehistoric (i.e. extinct before
1500) form from Maui in the Hawaiian Islands may represent a species or subspecies within this clade. The rate of
molecular evolution in
Haliaeetus is fairly slow, as is to be expected in long-lived birds which take years to successfully reproduce. In the
mtDNA cytochrome b gene, a
mutation rate of 0.5–0.7% per million years (if assuming an Early Miocene divergence) or maybe as little as 0.25–0.3% per million years (for a Late Eocene divergence) has been shown. The relationships to other genera in the family Accipitridae are less clear; they have long been considered closer to the genus
Milvus (kites) than to the true eagles in the genus
Aquila on the basis of their morphology and display behaviour; more recent genetic evidence agrees with this, but points to their being related to the genus
Buteo (buzzards/hawks), as well, a relationship not previously thought close. ==Description==