Evolution Esox is the one of three extant members of the family
Esocidae, alongside the physically dissimilar
Dallia and
Novumbra, the latter of which is its closest relative. Fossils of
Esox-like esocids are known as far back as the
Late Cretaceous in freshwater habitats of North America. Indeterminate
Esox fossils are known from the
Early Paleocene-aged
Fort Union Formation of Montana, US and the
Ravenscrag Formation of Saskatchewan, Canada, which represent the oldest records of the genus, only about 70,000 years after the
Cretaceous–Paleogene extinction event. These early
Esox coexisted with their older relative, the extinct
Estesesox, and suggest that
Esox either evolved very shortly after the extinction event, or already existed by the Late Cretaceous and immigrated to these localities shortly after the extinction event.
Species Currently, seven recognized species are placed in this genus, within two
subgenera:
Hybrids between
Esox masquinongy and
Esox lucius are well-known and referred to as the
tiger muskellunge.
Fossil species '' '', the earliest known pickerel The following fossil species are known:'' • †
Esox dispar Sytchevskaya, 1976 - Late Oligocene of Omsk Oblast, Russia • †
Esox kronneri Grande, 1999 -
Early Eocene of
Wyoming, US (
Green River Formation)'' • †
Esox moldavicus Sytchevskaya, 1974 - Pliocene to
Early Pleistocene of
Ukraine &
Moldova • †
Esox papyraceus Troschel, 1854 - Late Oligocene of Germany • †
Esox primaevus Gaudant, 1978 -
Early Oligocene of France • †
Esox nogaicus Kovalchuk, Wilson & Grande, 2017 - Early Pleistocene of Ukraine • †
Esox sibiricus Sytchevskaya, 1974 - Late Miocene and
early Pliocene of Ukraine, Russia (Omsk,
Tuva Republic,
Transbaikal), Kazakhstan & Mongolia • †
Esox tiemani Wilson, 1980 - Late Paleocene of Canada (
Paskapoo Formation) The oldest fossil species of
Esox is
Esox tiemani, from the late
Paleocene of Canada, which differs little from modern species. Two additional fossil species, both from the
Late Cretaceous of Alberta, Canada, are placed in their own genera:
Estesesox foxi (
Santonian to
Campanian), and
Oldmanesox canadensis).
Esox kronneri is one of the rarest known fish from the
Eocene-aged
Fossil Butte lagerstatte of Wyoming, USA.
E. kronneri appears to be more closely related to the North American pickerels (subgenus
Kenoza) than to the circumboreal "pike" (subgenus
Esox) lineage, suggesting these two lineages diverged by the
Early Eocene. The alleged
Late Cretaceous species "
Esox monasteriensis" von der Marck, 1863 is thought to be an indeterminate teleost. Other fossil fish genera such as
Enchodus and
Platinx also had their
type species be initially described under
Esox.'''' ==Diet==