Archaeogenetics has allowed the use of genetic analysis to trace migration patterns.
Kurgan/Steppe hypothesis from the
Pontic steppes and across Central Asia The Kurgan hypothesis, or steppe theory, is the most widely accepted proposal to identify the Proto-Indo-European homeland from which the Indo-European languages spread out throughout Europe and parts of Asia. It postulates that the people of a Kurgan culture in the
Pontic steppe north of the
Black Sea were the most likely speakers of the Proto-Indo-European language (PIE). The term is derived from the Russian (), meaning 'tumulus' or 'burial mound'.
R1a and R1b According to three
autosomal DNA studies,
haplogroups R1b and R1a, now the most common in Europe (R1a is also very common in
South Asia) would have expanded from the Pontic steppes, along with the Indo-European languages; they also detected an autosomal component present in modern Europeans which was not present in Neolithic Europeans, which would have been introduced with paternal lineages R1b and R1a, as well as Indo-European languages. Studies which analysed ancient human remains in
Ireland and
Portugal suggest that R1b was introduced in these places along with autosomal DNA from the Pontic steppes.
R1a and R1a1a The
subclade R1a1a (R-M17 or R-M198) is most commonly associated with Indo-European speakers. Data so far collected indicate that there are two widely separated areas of high frequency: • one in
Eastern Europe, around
Poland,
Ukraine, and
Russia, • the other in
Southern Asia, around the
Indo-Gangetic Plain, which extends to parts of
India,
Pakistan,
Bangladesh, and
Nepal. The historical and prehistoric possible reasons for this are the subject of on-going discussion and attention amongst population geneticists and genetic genealogists, and are considered to be of potential interest to linguists and archaeologists also. A large study in 2014 by Underhill et al., using 16,244 individuals from over 126 populations from across Eurasia, concluded there was compelling evidence that R1a-M420 originated in the vicinity of
Iran. The mutations that characterize haplogroup R1a occurred around 10,000 years
BP. Its defining mutation (M17) occurred about 10,000 to 14,000 years ago.
Yamnaya culture According to Jones et al. (2015) and ,
autosomal tests indicate that the Yamnaya-people were the result of admixture between "
Eastern Hunter-Gatherers" from eastern Europe (EHG) and "
Caucasus hunter-gatherers" (CHG).
Eastern European hunter-gatherers According to , "Eastern European hunter-gatherers" who inhabited Russia were a distinctive population of hunter-gatherers with high affinity to a ~24,000-year-old Siberian from the
Mal'ta-Buret' culture, or other,
closely related Ancient North Eurasian (ANE) people from Siberia and to the
Western Hunter-Gatherers (WHG). Their genomes showed that a continued mixture of the Caucasians with Middle Eastern took place up to 25,000 years ago, when the coldest period in the last Ice Age started. They concluded that about 4,500 years ago there was a major influx into Europe of
Yamnaya culture people originating from the
Pontic–Caspian steppe north of the Black Sea and that the DNA of
copper-age Europeans matched that of the Yamnaya.
Bronze Age Greeks A 2017
archaeogenetics study of Mycenaean and Minoan remains published in the journal
Nature concluded that the Mycenaean Greeks were genetically closely related with the
Minoans but unlike the Minoans also had a 13–18% genetic contribution from Bronze Age steppe populations.
Anatolian hypothesis Luigi Luca Cavalli-Sforza and
Alberto Piazza argue that Renfrew and Gimbutas reinforce rather than contradict each other. states that "It is clear that, genetically speaking, peoples of the Kurgan steppe descended at least in part from people of the Middle Eastern Neolithic who immigrated there from Turkey." state that:
Spencer Wells suggests in a 2001 study that the origin, distribution and age of the
R1a1 haplotype points to an ancient migration, possibly corresponding to the spread by the Kurgan people in their expansion across the
Eurasian steppe around 3000 BC. About his old teacher Cavalli-Sforza's proposal, states that "there is nothing to contradict this model, although the genetic patterns do not provide clear support either", and instead argues that the evidence is much stronger for Gimbutas' model:
Iranian/Armenian hypothesis David Reich (2018), noting the presence of some Indo-European languages (such as Hittite) in parts of ancient Anatolia, argues that "the most likely location of the population that first spoke an Indo-European language was south of the Caucasus Mountains, perhaps in present-day Iran or Armenia, because ancient DNA from people who lived there matches what we would expect for a source population both for the Yamnaya and for ancient Anatolians." Yet, Reich also notes that "...the evidence here is circumstantial as no ancient DNA from the Hittites themselves has yet been published."
Kristian Kristiansen, in an interview with
Der Spiegel in May 2018, stated that the Yamnaya culture may have had a predecessor at the Caucasus, where "proto-proto-Indo-European" was spoken. Recent DNA-research has led to renewed suggestions of a Caucasian homeland for the 'proto-Indo-Europeans'. According to Kroonen et al. (2018) and Damgaard et al. (2018), ancient Anatolia "show no indication of a large-scale intrusion of a steppe population." They further note that this lends support to the
Indo-Hittite hypothesis, according to which both proto-Anatolian and proto-Indo-European split-off from a common mother language "no later than the 4th millennium BCE." states that "the Armenian plateau hypothesis gains in plausibility" since the
Yamnaya partly descended from a Near Eastern population, which resembles present-day
Armenians." Wang et al. (2018) note that the Caucasus served as a corridor for gene flow between the steppe and cultures south of the Caucasus during the Eneolithic and the Bronze Age, stating that this "opens up the possibility of a homeland of PIE south of the Caucasus." However, Wang et al. also comment that the most recent genetic evidence supports an expansion of proto-Indo-Europeans through the steppe, noting: "but the latest ancient DNA results from South Asia also lend weight to a spread of Indo-European languages "via the steppe belt. The spread of some or all of the proto-Indo-European branches would have been possible via the North Caucasus and Pontic region and from there, along with pastoralist expansions, to the heart of Europe. This scenario finds support from the well attested and now widely documented '
steppe ancestry' in European populations, the postulate of increasingly patrilinear societies in the wake of these expansions (exemplified by R1a/R1b), as attested in the latest study on the Bell Beaker phenomenon."
David W. Anthony in a 2019 analysis, criticizes the "southern" or "Armenian" hypothesis (addressing Reich, Kristiansen, and Wang). Among his reasons being: that the Yamnaya lack evidence of genetic influence from the Bronze Age or late neolithic Caucasus (deriving instead from an earlier mixture of Eastern European hunter-gatherers and Caucasus hunter-gatherers) and have paternal lineages that seem to derive from the hunter-gatherers of the Eastern European Steppe rather than the Caucasus, as well as a scarcity in the Yamnaya of the Anatolian Farmer admixture that had become common and substantial in the Caucasus around 5,000 BC. Anthony instead suggests a genetic and linguistic origin of proto-Indo-Europeans (the Yamnaya) in the Eastern European steppe north of the Caucasus, from a mixture of these two groups (EHG and CHG). He suggests that the roots of Proto-Indo-European ("archaic" or proto-proto-Indo-European) were in the steppe rather than the south and that PIE formed mainly from a base of languages spoken by Eastern European hunter-gathers with some influences from languages of Caucasus hunter-gatherers. ==See also==