The placement of a cluster on the vine is similar to that of a
tendril, as both develop from the same uncommitted
primordia, the
anlagen. The grape bunch position on the side of the stem opposing a leaf is unusual for inflorescence of the plants. The typical shape of a cluster depends on the
grape variety. The bunch of grapes, like a tendril, has two
arms. The inner arm develops into a full-grown cluster, the smaller outer one might die off, develop into a small tendril-like arm with no fruit, or form a large "wing" with berries that sometimes ripen differently than the ones of the main cluster (for example, in
Trebbiano). The grape cluster is shaped as a
panicle, with the main branch ("axis") called
rachis. Additional branches ("shoulders" or
branch-twigs) are connected to the rachis irregularly and split into
pedicels (short stems attaching individual berries). In a large cluster, the shoulders might bear secondary (
sub-twigs) and sometimes tertiary branches before ending up with pedicels (the twig carrying the pedicels is also known as a
terminal pedicel twig). The beginning of the rachis before the first split is called a
peduncle. In a ripe cluster the peduncle might be the only fully visible part of the cluster branches. The arrangement of the branches, so called
cluster architecture, determines the distribution of berries and free space within the cluster. The architecture is complex and influences yield, quality, and resistance to pests and diseases. While mostly determined by the
ramification patterns of the rachis, its development can be influenced by management technique, like
leaf removal. A
relational growth grammar (RGG) can be used to describe the architecture of a cluster. == Development ==