They were originally classified as the subfamily Geotrupinae in the family Scarabaeidae before being elevated to a family. Traditionally the family
Bolboceratidae was included (as the subfamily Bolboceratinae) on the basis of the number of
antenna segments, but examination of a different set of characteristics prompted Scholtz & Browne (1995) to elevate Bolboceratidae to a family, a result supported by recent phylogenetic research. The family has more than 600 species in about 30 genera in two subfamilies; more recent phylogenetic studies indicate that Taurocerastinae is not related to Geotrupinae, and is instead more closely related to
Lucanidae and
Diphyllostomatidae. •
Geotrupinae •
Allotrypes François, 1904 (= "
Allotrupes" Boucomont, 1912) •
Anoplotrupes Jekel, 1866 •
Baraudia López-Colón, 1996 •
Ceratophyus Fischer von Waldheim, 1823 •
Ceratotrupes Jekel, 1865 •
Chelotrupes Jekel, 1866 •
Cnemotrupes Jekel, 1866 •
Cretogeotrupes Nikolajev, 1992 •
Enoplotrupes Lucas, 1869 •
Geohowdenius Zunino, 1984 •
Geotrupes Latreille, 1796 •
Halffterius Zunino, 1984 •
Haplogeotrupes Nikolaev, 1979 •
Jekelius López-Colón, 1989 •
Lethrus Scopoli, 1777 •
Megatrupes Zunino, 1984 •
Mycotrupes LeConte, 1866 •
Odontotrypes Fairmaire, 1887 •
Onthotrupes Howden, 1964 •
Phelotrupes Jekel, 1866 •
Peltotrupes Blanchard, 1888 •
Pseudotrypocopris Miksic, 1954 •
Sericotrupes Zunino, 1984 •
Silphotrupes Jekel, 1866 •
Thorectes Mulsant, 1842 •
Trypocopris Motschulsky, 1860 •
Typhaeus Leach, 1815 •
Zuninoeus López-Colón, 1989 •
Taurocerastinae •
Frickius Germain, 1897 •
Taurocerastes Philippi, 1866 ==References==