Kitadani Formation

The Kitadani Formation is a unit within the Tetori Group, a Lower Cretaceous sequence of predominantly sedimentary rock which crops out in the Fukui, Ishikawa, and Gifu prefectures of west-central Honshu, Japan in the region surrounding Mount Haku. The formations present within the Tetori Group vary laterally, and the Kitadani Formation crops out only in the Kuzuryū River district of Fukui Prefecture. In this region, the sequence comprises, in ascending stratigraphic order: Gomijima Formation, the Kuwajima Formation, the Akaiwa Formation, and Kitadani Formation. The Kitadani Formation has had varying nomenclature throughout the history of its study. the "Chinaboradani Alternation of Tuff, Shale, and Sandstone", the "Kitadani Alternation of Sandstone, Shale, and Tuff", and simply the "Kitadani Alternation" prior to its designation as a formation. == Age ==

The Kitadani Formation was biostratigraphically dated to the late Barremian and early Aptian ages of the Early Cretaceous Epoch in 2002 based upon the presence of the freshwater bivalve Nippononaia ryosekiana. In 2005, part of the Kitadani Formation was biostratigraphically dated to the Barremian Age based upon the occurrence of the charophyte gyrogonite Clavator harrisii reyi in association with other charophytes. These biostratigraphic age assignments are supported by zircon fission track radioisotopic ages of tuff, which date the Kitadani Formation to 127-115 Ma. However, the age of the overlying lower Akaiwa Formation was estimated around 121.2±1.1 Ma based on U-Pb zircon dating, so the age of the Kitadani Formation is more likely to be late Aptian. ==Fossil assemblage==

The Kitadani Formation preserves a diverse assemblage of plant fossils; invertebrate fossils; and vertebrate body and trace fossils, including mammals, turtles, crocodylomorphs, and dinosaurs. Many vertebrate specimens from the Kitadani Formation are incomplete and poorly preserved, so taxonomic diversity is likely higher than it seems. Plant fossils The plant fossil assemblage of the Kitadani Formation is characterized by a rarity of ferns and an abundance of cycadales and conifers represented mostly by cones and shoots. A palynological study in 2013 resulted in the identification of greater than 40 species of spores, pollen grains, and plant fragments from the Kitadani Formation representing gymnosperms, freshwater algae, and epiphyllous fungus; however, no angiosperm pollen was identified. Branches of the conifer Brachyphyllum obesum have been recovered, which was interpreted to represent the warming and possible drying of the climate toward the upper Tetori Group. This interpretation is supported by the lack of plants from lower in the Tetori Group, such as ginkgos, in the Kitadani Formation. Invertebrate fossils The invertebrate fossil assemblage of the Kitadani Formation mostly comprises freshwater and brackish water bivalve and gastropod mollusks. Vertebrate fossils Actinopterygii Mammaliaformes Testudines Turtles are represented mostly by shell fragments within the Kitadani Formation. Crocodylomorpha A nearly complete skeleton of goniopholidid has been discovered from the Kitadani Formation, but this material remains undescribed. Dinosauria Dinosaurs are among the most well-known vertebrate taxa from the Kitadani Formation. Taxa from all three major dinosaurian clades — Theropoda, Sauropodomorpha, and Ornithischia — have been recovered. Ichnofossils Undescribed tracks of ankylosaurs and sauropods have been discovered from the Kitadani Formation. Eggs == See also ==