Veracruz moist forests

The Veracruz moist forests cover an area of , occupying a portion of Mexico's Gulf Coastal Plain between the Sierra Madre Oriental and the Gulf of Mexico. The forests extend from southern Tamaulipas across northern Veracruz, eastern San Luis Potosí, and portions of eastern Hidalgo, northeastern Puebla and northern Queretaro. The Huasteca region includes much of the ecoregion. To the north, the forests transition to the dry lowland Tamaulipan mezquital and the upland Tamaulipan matorral. To the west, the Sierra Madre Oriental pine–oak forests occupy the higher elevations of the Sierra Madre Oriental. The Moctezuma River and its tributaries have carved deep canyons through the Sierra Madre, which allow moist air from the Gulf of Mexico to flow further west into the plateaus and mountains, including the Sierra Gorda, and the moist forests extend westwards along the river valleys. South of the gap where the Moctezuma River cuts through the Sierra Madre, the Veracruz montane forests and Oaxacan montane forests occupy middle slopes of the Sierra. The Veracruz dry forests separate the Veracruz moist forests from the Petén–Veracruz moist forests further south. The northernmost extension of the Veracruz moist forests occurs in the El Cielo Biosphere and the Sierra de Tamaulipas at a latitude of about 23° 20′ degrees north. ==Climate==

The climate of the region is tropical and humid, with rains during seven months of the year and mild variation in temperature. Average annual rainfall is . == Gallery ==

File:Rio Guayalejo & Cerro del Bernal, Municipality of Gonzlaez, Tamaulipas, Mexico (17 April 2001).jpg|Rio Guayalejo and Cerro del Bernal, Municipality of González, Tamaulipas (17 April 2001) File:Rio Sabinas, Municipality of Gómez Farías, Tamaulipas, Mexico (15 April 2001).jpg|Río Sabinas, Municipality of Gómez Farías, Tamaulipas (15 April 2001). File:Estero del Rio Tuxpan.jpg|Rio Tuxpan (Zona Juana Moza), Veracruz, Mexico File:Cascada en Tlaxcalantongo Puebla - panoramio.jpg|Forest near Tlaxcalantongo, municipality of Xicotepec, Puebla (16 March 2009). File:En Coatepec, Veracruz (28 June 2008).jpg|In Coatepec, Veracruz (28 June 2008). ==Flora==

The canopy of this ecoregion is characterized by trees reaching a height of up to , such as Mayan breadnut (Brosimum alicastrum), sapodilla (Manilkara zapota), rosadillo (Celtis monoica), Bursera simaruba, Dendropanax arboreus, and Sideroxylon capiri. The southern parts of the ecoregion feature mahogany (Swietenia macrophylla), Manilkara zapota, Bernoullia flammea, and Astronium graveolens. ==Fauna==

Mammals: Three species of rodents are endemic to this area, the El Carrizo deer mouse (Peromyscus ochraventer) the Tamaulipan woodrat (Neotoma angustapalata), and the Jico crested tail mouse (Habromys simulatus). Spider monkeys (Ateles geoffroyi), the northernmost representative of the New World primates range into this region, although it is an endangered species and not common. Marsupials include the common opossum (Didelphis marsupialis) and Mexican mouse opossum (Marmosa mexicana). Other mammals such as the Mexican anteater (Tamandua mexicana), lowland paca (Cuniculus paca), and red brocket (Mazama americana) are known from this region. Among the carnivores are the kinkajou (Potos flavus), mustelids such as tayra (Eira barbara) and greater grison (Galictis vittata), and five cat species including jaguarondi (Herpailurus yaguarondi), ocelot (Leopardus pardalis), margay (Leopardus wiedii), Puma (Puma concolor) and jaguar (Panthera onca). Just a few of the many species of bats include the elegant myotis (Myotis elegans), wrinkle-faced bat (Centurio senex), and hairy-legged vampire bat (Diphylla ecaudata). File:Mono Araña Centroamericano - panoramio.jpg|Spider monkey (Ateles geoffroyi), near Catemaco, Veracruz (4 March 2015). Birds: Endemic birds include the red-crowned amazon (Amazona viridigenalis), Altamira yellowthroat (Geothlypis flavovelata), and crimson-collared grosbeak (Rhodothraupis celaeno). File:Garza tigre mexicana - panoramio.jpg|Bare-throated tiger heron (Tigrisoma mexicanum), Veracruz, Mex. (15 July 2011). File:Boat-billed heron (Cochleatius cochlearius), Campeche, Mex. (2015).jpg|Boat-billed heron (Cochleatius cochlearius), Campeche, Mex. (2015). File:Collared Forest Falcon (Micrastur semitorquatus), Campeche, México (14 December 2015).jpg|Collared forest falcon (Micrastur semitorquaatus), Campeche, Mex. (2015). File:Blue-crowned motmot (Momotus momota), Ixhuatán, Chiapas (8 March 2016).jpg|Lesson's motmot (Momotus lessonii), Chiapas, Mex. (2016). File:Lineated woodpecker (Dryocopus lineatus), Laguna de Sontecomapan, Veracruz (4 January 2014).jpg|Lineated woodpecker (Dryocopus lineatus), Veracruz, Mex. (2014). File:Ivory-billed woodcreeper (Xiphorhynchus flavigaster), Catemaco, Veracruz (10 January 2014).jpg|Ivory-billed woodcreeper (Xiphorhynchus flavigaster), Veracruz, Mex. (2014). File:Megarynchus pitangua San Juan Evangelista.jpg|Boat-billed flycatcher (Megraynchus pitangua), Veracruz, Mex. (2011). File:Yellow-throated euphonia (Euphonia hirundinacea), Laguna de Sontecomapan, Veracruz (4 January 2014).jpg|Yellow-throated euphonia (Euphonia hirundinacea), Veracruz, Mex. (2014). File:Scrub euphonia (Euphonia affinis), Matlapa, San Luis Potosi, Mexico (11 March 2014).jpg|Scrub euphonia (Euphonia affinis), San Luis Potosí, Mex. (2014). Reptiles: The Morelet's crocodile (Crocodylus moreletii) can be found in the remote areas of the rivers and lagoons in this region with turtles like the Mesoamerican slider (Trachemys venusta) and scorpion mud turtles (Kinosternon scorpioides). Herrera's mud turtle (Kinosternon herrerai) and the Mexican box turtle (Terrapene mexicana) are endemic to this region. Endemic lizards include the rare plain-necked glass lizard (Ophisaurus incomptus) and the Cave Tropical Night Lizard (Lepidophyma micropholis) known only from caves and the vicinity of cave openings in the Sierra Cucharas/Sierra del Abra of southern Tamaulipas adjacent San Luis Potosí. Other lizards found in the region include the silky anole (Anolis sericeus), rainbow ameiva (Holcosus undulatus), rose-belly lizard (Sceloporus variabilis), Mexican spiny-tailed iguana (Ctenosaura acanthura), and casque-headed lizard (Laemanctus serratus). Snakes such as Iverson's threadsnake (Rena iversoni), the brown hook-nose snake (Ficimia olivacea), Taylor's cantil (Agkistrodon taylori), and the Totonacan rattlesnake (Crotalus totonacus) are largely associated with the Veracruz moist forest but all range into areas beyond the strict limits of this region. Many tropical snakes from Central America range into this region like the Central American boa constrictor (Boa imperator), Central American indigo snake (Drymarchon melanurus), blunthead tree snake (Imantodes cenchoa), Mexican parrot snake (Leptophis mexicanus), brown vine snake (Oxybelis aeneus), tropical ratsnake (Pseudelaphe flavirufa), tropical tree snake (Spilotes pullatus), green ratsnake (Senticolis triaspis), banded snail sucker (Tropidodipsas fasciata), and terrestrial snail sucker (Tropidodipsas sartorii). In addition to the cantil and Totonacan rattlesnake, venomous snakes from this province include the Texas coral snake (Micrurus tener) and the terciopelo (Bothrops asper). File:Terrapene mexicana, Mexican Box Turtle, Tamaulipas.jpg|Mexican box turtle (Terrapene mexicana), southern Tamaulipas (30 May 2005). File:Kinosternon scorpioides Scorpion Mud Turtle, Tamaulipas.jpg|Scorpion mud turtle (Kinosternon scorpioides), southern Tamaulipas (23 September 2004). File:Ctenosaura acanthura, Mexican Spiny-tailed Iguana, Tamaulipas.jpg|Mexican spiny-tailed iguana (Ctenosaura acanthura) Municipality of Ocampo, Tamaulipas (13 September 2007). File:Crocodylus moreletii, Morelet’s Crocodile, Laguna del Carppintero.jpg|Morelet's crocodile (Crocodylus moreletii), Laguna del Carppintero, Tamaulipas (21 May 2007). File:Boa imperator, Central American Boa Constrictor Tamaulipas.jpg|Central American boa constrictor (Boa imperator), Municipality of Gómez Farías, Tamaulipas (23 August 2007). File:Agkistrodon taylori, Taylor’s Cantil, Tamaulipas.jpg|Taylor's cantil (Agkistrodon taylori) southern Tamaulipas (24 October 2004). File:Crotalus totonacus, Totonacan Rattlesnake, Tamaulipas.jpg|Totonacan rattlesnake (Crotalus totonacus) southern Tamaulipas (10 July 2005). Amphibians: Although salamander diversity in Mexico is among the highest in the world, they mostly occur in mountainous areas. The broadfoot mushroomtongue salamander (Bolitoglossa platydactyla), a lungless salamander of the Plethodontidae family, occupies the coastal lowlands from sea level up to about 1000 meters from southern Tamaulipas to the Isthmus of Tehuantepec. The distribution of the southern subspecies of the black-spotted newt (Notophthalmus meridionalis kallerti) nearly matches that of the Veracruz moist forests region. There are also enigmatic reports of an isolated population of the lesser siren (Siren intermedia) from central Veracruz. Anuran, or frog diversity in the region is higher and among the species found in this area are the Gulf Coast toad (Incilius nebulifer), cane toad (Rhinella horribilis), white-lipped frog (Leptodactylus fragilis), sabinal frog (Leptodactylus melanonotus), elegant narrow-mouthed toad (Gastrophryne elegans), sheep frog (Hypopachus variolosus), and the burrowing toad (Rhinophrynus dorsalis). The veined treefrog (Trachycephalus typhonius) is a large species that produces sticky, mucous, skin secretions, presumed to deter desiccation in the dry season, as well as being a toxin to predators. One source list 93 species in the Guayalejo-Temesí watershed of Tamaulipas alone, but this includes brackish-marine species. Many of the streams and rivers of this region are spring fed by karstic aquifers, providing consistent and relatively thermally stable water compared to other basins sourced largely by precipitation. The karstic environment of the Sierra Madre Oriental produces many caves and subterranean waterways that ultimately surface near the base of the mountains in the west of this region. Some fish like the endemic phantom blindcat (Prietella lundbergi) are adapted to caves and have been collected at depths of 50 meters in cave systems of the Rio Frio (in the Rio Guayalejo drainage). Some populations of the Mexican tetra (Astyanax mexicanus [= A. jordani]) also inhabit caves and are blind, although other populations of the same species living in surface streams have eyes, while still other populations are intermediate. The region is rich in platyfish and swordtail (Xiphophorus) diversity including sheepshead swordtail (Xiphophorus birchmanni), delicate swordtail (Xiphophorus cortezi), mountain swordtail (Xiphophorus nezahualcoyotl), and the endemic variable platyfish (Xiphophorus variatus). Cichlid diversity is also high in this area including the lowland cichlid (Herichthys carpintis) and endemic species like the chairel cichlid (Herichthys pantostictus|Herichthys [Nosferatu] pantostictus) and Nautla cichlid (Herichthys deppii) among others. Other fishes include the pigmy shiner (Notropis tropicus), Forlón gambusia (Gambusia regani), gulf gambusia (Gambusia vittata), chubsucker minnow (Dionda erimyzonops), and the endemic lantern minnow (Dionda ipni). Conservation threats of fishes include, damming of waterways, water diversion for agriculture, runoff associated with agriculture and livestock, oil and industrial contamination, invasive species, and irresponsible recreational activities. File:Bolitoglossa platydactyla, Broadfoot Mushroomtongue Salamander, Tamaulipas.jpg|Broadfoot mushroomtongue salamander (Bolitoglossa platydactyla), municipality of Ocampo, Tamaulipas (11 July 2005). File:Leptodactylus melanonotus WLF-255-7A, Municipio Aldama; 3 June 2004.jpg|Sabinal frog (Leptodactylus melanonotus), municipality of Aldama, Tamaulipas (3 June 2004). File:Smilisca baudinii, Mexican Treefrog, Tamaulipas.jpg|Mexican treefrog (Smilisca baudinii), municipality of Gómez Farías, Tamaulipas (8 August 2004). File:Trachycephalus typhonius, Veined Treefrog, Tamaulipas.jpg|Veined treefrog (Trachycephalus typhonius), municipality of Gómez Farías, Tamaulipas (5 June 2002). File:Epigean and cave-dwelling Mexican tetras Astyanax mexicanus.jpg|Epigean and cave-dwelling Mexican tetras (Astyanax mexicanus). File:Xipho variatus maschio.jpg|Variable platyfish (Xiphophorus variatus), male. File:Herichthys carpintis 01.jpg|Lowland cichlid (Herichthys carpintis). ==Conservation and threats==

The forests have been heavily altered by human activity, so that only a few enclaves of mature forest remain. Forests have been cleared for timber harvesting, agriculture, and grazing, and much of the original forest has been replaced with scrubland or secondary forest. ==Protected areas==

7.66% of the ecoregion is in protected areas. Protected areas entirely or partly within the ecoregion include El Cielo, Sierra del Abra Tanchipa, Sierra Gorda, Sierra Gorda de Guanajuato, and Sierra de Tamaulipas biosphere reserves, Laguna Madre and Río Bravo Delta Flora and Fauna Protection Area, Bernal de Horcasitas, El Sótano de Las Golondrinas, and La Hoya de las Huahuas natural monuments, Río Filo-Bobos y su Entorno protected area, and Zona Protectora Forestal Vedada Cuenca Hidrográfica del Río Necaxa natural resources protection area. ==See also==