Gempylidae

• Subfamily Clade 1 Mthethwa, 2023 • Genus Diplospinus • Diplospinus multistriatus Maul, 1948 (Striped escolar) • Genus Gempylus • Gempylus serpens G. Cuvier, 1829 (Snake mackerel) • Genus Nealotus • Nealotus tripes J. Y. Johnson, 1865 (Black Snake mackerel) • Genus Nesiarchus • Nesiarchus nasutus J. Y. Johnson, 1862 (Black gemfish) • Genus Paradiplospinus • Paradiplospinus antarcticus Andriashev, 1960 (Antarctic escolar) • Paradiplospinus gracilis (A. B. Brauer, 1906) (Slender escolar) • Genus Promethichthys • Promethichthys prometheus (G. Cuvier, 1832) (Roudi escolar) • Genus Rexea • Rexea alisae C. D. Roberts & A. L. Stewart, 1997 • Rexea antefurcata Parin, 1989 (Long-finned escolar) • Rexea bengalensis (Alcock, 1894) (Bengal escolar) • Rexea brevilineata Parin, 1989 (Short-lined escolar) • Rexea nakamurai Parin, 1989 (Nakamura's escolar) • Rexea prometheoides (Bleeker, 1856) (Royal escolar) • Rexea solandri (G. Cuvier, 1832) (Silver gemfish) • Genus Rexichthys • Rexichthys johnpaxtoni Parin & Astakhov, 1987 (Paxton's escolar) • Genus Leionura • Leionura atun (Euphrasén, 1791) (Snoek) • Genus Thyrsites • Thyrsites lepidopodea (Lesson, 1831) (White snake mackerel)) • Genus Thyrsitoides • Thyrsitoides marleyi Fowler, 1929 (Blacksail snake mackerel) • Thyrsitoides zarahoustrae Arambourg, 1967 • Genus Tongaichthys • Tongaichthys robustus I. Nakamura & E. Fujii, 1983 (Tonga escolar) • Subfamily Clade 2 Mthethwa, 2023 • Genus Epinnula • Epinnula magistralis Poey, 1854 • Epinnula pacifica Ho, Motomura, Hata & Jiang, 2017 • Genus Lepidocybium • Lepidocybium flavobrunneum (A. Smith, 1843) (Escolar) • Genus Neoepinnula • Neoepinnula americana (M. G. Grey, 1953) (American sackfish) • Neoepinnula minetomai Nakayama, Y. Kimura & Endo, 2014 (Large-eyed sackfish) • Neoepinnula orientalis (Gilchrist & von Bonde, 1924) (Sackfish) • Genus Ruvettus • Ruvettus pretiosus Cocco, 1833 (Oilfish) The Gempylidae are broadly categorized into two clades; Clade 1, which includes more derived & elongate genera, and Clade 2, which includes more basal & fusiform genera. The Trichiuridae are an outgroup. }} Gempylidae are believed to have first evolved at least 20 million years after the Late Cretaceus Extinction event, potentially due to tectonic plate movements. • †Abadzekhia Bannikov, 1985 (Early Oligocene of North Caucasus, Russia and Germany) • †Contemptor Calzoni, Giusberti & Carnevale, 2026 (Eocene of northern Italy) • †Chelifichthys Carnevale, 2006 (Late Miocene (Messinian) of Algeria) • †Eothyrsites Chapman, 1934 (mid-late Eocene of New Zealand) • †Hemithyrsites Daniltshenko, 1960 (Oligocene of Poland & Romania) • †Krampusichthys Calzoni, Giusberti & Carnevale, 2025 (Early Eocene of Italy) • †Laurinichthys Calzoni, Giusberti & Carnevale, 2025 (Early Eocene of Italy) • †Progempylus Casier, 1966 (Early Eocene of England) • †Thyrsitocephalus vom Rath, 1859 (Early Oligocene of Switzerland) • †Wudelenia Calzoni, Giusberti & Carnevale, 2025 (Early Eocene of Italy) ==Timeline==