The birds formerly considered to be the resident
Indian
subspecies are now considered a separate
species, the
Indian spotted eagle (
Clanga hastata), quite distinct and readily separable by
morphological,
behavioral,
ecological, and
DNA sequence data. The European
taxon is actually closer to the greater spotted eagle; their common ancestor seems to have diverged around the middle
Pliocene, perhaps some 3.6 million years ago, from the ancestors of the Indian birds. The "proto-spotted eagle" probably lived in the general region of
Afghanistan, being split into a northern and a southern lineage when both
glaciers and
deserts advanced in Central Asia as the
last ice age began. The northern lineage subsequently separated into the eastern (greater) and western (lesser) species of today, probably around the Pliocene-
Pleistocene boundary not quite 2 million years ago. The present species
hybridizes occasionally with the greater spotted eagle. Hybrid birds are almost impossible to identify if not seen up close. ==Distribution and habitat==