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Tuta absoluta

Tuta absoluta or Phthorimaea absoluta is a species of moth in family Gelechiidae known by the common names South American tomato pinworm, tomato leafminer, tomato pinworm and South American tomato moth. It is well known as a serious pest of tomato crops in Europe, Africa, western Asia and South and Central America, with larvae causing up to 100% loss if not effectively controlled.

Naming history
T. absoluta was originally described in 1917 by Edward Meyrick as Phthorimaea absoluta, based on individuals collected from Huancayo (Peru). Scrobipalpula absoluta (Povolný), ==Biology==
Biology
The larva feeds voraciously upon tomato plants, producing large galleries in leaves, burrowing in stalks, and consuming apical buds and green and ripe fruits. It is capable of causing a yield loss of 100%. Morphology Adults are in length and present filiform antennae and silver to grey scales. Black spots are present on anterior wings, and the females are wider and more voluminous than the males. The adult moth has a wingspan around . In favorable weather conditions, eight to ten generations can occur in a single year. Hosts Tomato is the main host plant, but T. absoluta also attacks other crop plants of the nightshade family, including potato, This introduction of other hosts is due to multiple relocations of the agriculture of these crops. It is known from many solanaceous weeds, including Datura stramonium and Solanum nigrum. Also known from non-Solanaceae hosts in the Amaranthaceae, Convolvulaceae, Fabaceae, and Malvaceae. Laboratory rearing Laboratory rearing is abnormally difficult because T. absoluta requires maternal leaf contact with a suitable host plant for oviposition. ==Global spread==
Global spread
This moth was first known as a tomato pest in many South American countries (and Easter Island) and South Africa in 2016. Although it is not there yet, researchers at the University of Guam are concerned about the possible spread of T. absoluta to Guam. USDA's Animal and Plant Health Inspection Service assumed T. absoluta to be present in most of sub-Saharan Africa. since the 2010-11 survey. of their own and neighboring countries (including India and Pakistan) that already have the pest. Surveillance occurs in production areas and near international airports. ==Damage==
Damage
Losses on tomatoes can reach 100% due to larval feeding, if not effectively controlled. Even if not that severe, damage will require postharvest inspection expenditures and some financial loss due to unattractive fruit. The initial European invasion increased tomato production costs by more than 450/hectare. ==Management==
Management
Some populations of T. absoluta have developed resistance to organophosphate and pyrethroid pesticides. Newer compounds such as spinosad, imidacloprid, and Bacillus thuringiensis have demonstrated some efficacy in controlling European outbreaks of this moth. Insecticide costs have increased rapidly, and even that has not always produced good results, due to high quantity application of insecticides that are not especially effective against T. absoluta. As a result, new registrations have been obtained specifically for this pest starting in 2009. Between 2009 and 2011 there was a dramatic increase in authorized APIs and MoAs in Spain and Tunisia for this reason. Bacillus thuringiensis,|leftThe sex pheromone for T. absoluta has been identified by researchers at Cornell University and has been found to be highly attractive to male moths. Pheromone lures are used extensively throughout Europe, South America, North Africa and the Middle East for the monitoring and mass-trapping of T. absoluta. The use of pheromone products in combination with a yellow delta trap has been recorded in South Africa. This concept is used to monitor populations of T. absoluta in tomato orchards. Also the use of electric mosquito traps give good results. Insecticide resistance History and genetics Organophosphate and pyrethroid resistance developed in Chile, then in Brazil and (as noted above) Argentina. (The use of chlorantraniliprole for T. absoluta has also resulted in resistance in B. tabaci, even though it is not used against that species, merely because they co-occur on tomato. This is expected to make cyantraniliprole unusable if needed on B. tabaci, in the same area.) Voltage-dependent sodium channel blocker resistance Resistance to indoxacarb (IRAC group 22A) has appeared due to the mutations F1845Y and V1848I, but is not yet reported for another voltage-dependent sodium channel blocker, metafumizone (22B). (These two mutations, as with the diamides above, have P. xylostella analogues, but in this case these analogues are known to be effective against both indoxacarb and metafumizone.) Nicotinic acetylcholine receptor channel blocker resistance Cartap, a nicotinic acetylcholine receptor channel blocker (IRAC group 14), began to show low to moderate efficacy decline in South America starting in 2000, and increasing through at least 2016. Some of this is due to elevated cytochrome P450 activity (see below) possibly as part of demethylation and sulfoxidation detoxification, while less is thought to be due to esterases and glutathione S-transferases. (The use of cartap for T. absoluta has also resulted in resistance in B. tabaci, even though it is not used on that species, merely because they co-occur on tomato.) Cytochrome P450 and resistance Cytochrome P450s are used to resist: • cartap, :*possibly as part of demethylation and sulfoxidation detoxification, • possibly also abamectin :*along with resistance due to higher esterase activity, • but not pyrethroids, :*because although important in other insects, :*they are of limited usefulness in this case for reasons still unknown, but overall, specific information is still lacking connecting which particular P450s and which particular resistances. ==References==
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