The predominant plant community is
grassland, ranging from subalpine to alpine. Open forest of
Hartweg's pine (
Pinus hartwegii) and/or
Montezuma pine (
Pinus montezumae) marks the transition between the lower-elevation
pine-oak forests and the subalpine grasslands at about 3500 meters elevation. Grassland dominated by
Toluca fescue (
Festuca tolucensis) and
Toluca reedgrass (
Calamagrostis tolucensis) grows as an understory among the pines, in open
Juniperus monticola woodland, or as open grassland. Other grasses and forbs include
Arenaria bryoides, Descurainia impatiens, Draba jorullensis, Lupinus montanus, Muhlenbergia pusilla, M. quadridentata, Oxalis sp.,
Pedicularis orizabae, Penstemon gentianoides, Poa annua, Potentilla staminea, Sicyos parviflorus, and
Trisetum spicatum. At 3800 meters elevation
Muhlenbergia quadridentata becomes the dominant grass, with
Calamagrostis tolucensis dominant between 3800 and 4200 meters. The transition from subalpine to alpine grasslands occurs between 4200 and 4300 meters elevation. Alpine grasslands are dominated by the grasses
Festuca livida and
Arenaria bryoides. Other alpine zone plants include
Festuca tolucensis and species of
Carex, Cerastium, Cirsium, Draba, Eryngium, Gnaphalium, Lupinus, Luzula, Oxylobus, Phacelia, Plantago, Potentilla, Ranunculus, and
Senecio, together with ferns, mosses, and lichens. The flora of the Zacatonal is distinct from both temperate North America's alpine grasslands and the alpine
páramo of Central and South America, and includes many endemic species. ==Fauna==