Sheltopusik

The sheltopusik was previously included in the genus Ophisaurus, but has since been placed in its own genus Pseudopus. It was originally described in 1775 by Peter Simon Pallas as Lacerta apoda. There are three subspecies: • P. a. apodus (Pallas, 1775) – the type subspecies, ranging from Crimea through Transcaucasia, east to Central Asia as far as Kazakhstan • P. a. levantinus Jablonski, Ribeiro-Junior, Meiri, Maza, Mikulíček & Janzik, 2021 – restricted to the Levant, from southern Turkey south to Israel and Palestine • P. a. thracicus (Obst, 1978) – the westernmost subspecies, ranging from Croatia to Greece, east to western Turkey Despite only being described in 2021, P. a. levantinus is the largest and most genetically diverse of the subspecies, indicating an older and more complex evolutionary history compared to its sister subspecies. == Skull anatomy ==

The skull is elongated, twice as long as it is wide and widest at the posteroventral portions of the jugals, where the cheeks proceed to narrow into the snout. The exonarial fenestra (external naris) is similarly elongated and elliptical. The orbits, which lie in a posterior portion of the anterior part of the skull, are large. The premaxillary–maxilla has an elongated opening and has rounded ends. It is positioned between the forked premaxillary process of the maxilla and the maxillary and vomeromaxillary processes of the premaxilla. ==Description==

The sheltopusik can reach a length of . It is tan colored, paler on the ventral surface and the head, with a ring-like/segmented appearance that makes it look like a large earthworm with a distinctive fold of skin down each side called a lateral groove. Small (2-mm) rear legs are sometimes visible near the cloaca. Though the legs are barely discernible, the sheltopusik can be quickly distinguished from a snake by its ears, eyelids, and ventral scales. This species exhibit sexual dimorphism as males are on average larger than females. ==Habitat and behaviour==

P. apodus inhabits open country, such as short grassland or sparsely wooded hills. A preference for microhabitats that exhibit areas with sunlight and shade close together has been observed. It consumes arthropods and small mammals. Snails and slugs appear to be its favorite prey, which may explain why it is particularly active in wet weather, although it prefers a dry habitat. Breaking through the shells of snails is an especially easy task due to their teeth and jaw structure. ==Defensive behaviour==

their tails. Due to its size, the sheltopusik tends to respond to harassment by hissing, biting, and musking. It is less likely to drop off its tail than some other species that display caudal autotomy. However, these occasional displays of caudal autotomy are responsible for the name "glass lizard" (or "glass snake"). The released tail may break into pieces, leading to the myth that the lizard can shatter like glass and reassemble itself later. In reality, if the tail is lost, it grows back slowly, but is shorter and darker. The replacement tail may grow back to full length after an extended period of time. ==Reproduction==

About 10 weeks after mating, the female P. apodus lays about eight eggs, which she hides under bark or a stone, and often guards them. The young hatch after 45 to 55 days. They are typically about long and usually start to eat after four days. == In captivity ==

at the Artis Royal Zoo Sheltopusiks are frequently available in the exotic pet trade, though rarely captive-bred. They do not typically tolerate a large amount of handling, but they adapt to captivity well, feeding on crickets, meal worms, small mice, eggs, snails, or pieces of meat. They are even known to accept these meals from a keeper's tweezers, or even from their hands once they become used to captivity. However, sheltopusiks do get excited around food and have surprisingly powerful jaws. They make hardy captives, capable of living up to 50 years. ==Relationship with humans==

Remains of the Levant subspecies (P. a. levantinus) are known from Natufian sites in Israel, suggesting that it was eaten by the local population at the time. == Gallery ==

File:Pseudopus apodus, Uzbekistan, Surxondaryo Region, Kugitang Mountain Range imported from iNaturalist photo 224163004.jpg|In Uzbekistan File:Pseudopus apodus, West Greece, Peloponnese, Western Greece and the Ionian Islands, GR imported from iNaturalist photo 282395812.jpg|In Greece File:Pseudopus apodus, Meghri, Syunik Province, Armenia imported from iNaturalist photo 263273448.jpg|In Armenia File:Pseudopus apodus, Primorsko, Bulgaria imported from iNaturalist photo 213646243.jpg|In Bulgaria File:Circaetus gallicus with Pseudopus apodus, Εθνικό Δρυμό Πάρνηθας, Κρυονερίου, Восточная Аттика, GR imported from iNaturalist photo 207074472.jpg|Caught by a short-toed snake eagle File:Pseudopus apodus crawling.webm|Moving on a brick road ==References==