Pachygenelus had both an
articular-
quadrate and
dentary-
squamosal jaw joint characteristic of ictidosaurs. Only
mammals possess the dentary-squamosal articulation, while all other tetrapods possess the typical arcticular-quadrate articulation. Thus the jaw of
Pachygenelus can be seen as transitional between non-mammalian synapsids and true mammals. Another feature of
Pachygenelus that is shared with mammals is plesiomorphic prismatic enamel, or
enamel arranged into strengthened prisms. The upper and lower tooth rows
occluded with one another, although not as close as what is seen in true mammals. Wear facets are present on the lingual sides of the upper and external faces of the lower postcanines and are seen as evidence for the occlusion. Despite all of the derived adaptation seen in its teeth, the dentition of
Pachygenelus is believed to have been the least specialized of all tritheledontids. == Taxonomy ==