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Panamanian golden frog

The Panamanian golden frog, also known as the Panamanian golden toad, Zetek's golden frog, and Cerro Campana stubfoot toad and other names, is a species of toad endemic to Panama. Panamanian golden frogs inhabit the streams along the mountainous slopes of the Cordilleran cloud forests of west-central Panama. While the IUCN lists it as Critically Endangered, it may in fact have been extinct in the wild since 2007. Individuals have been collected for breeding in captivity in a bid to preserve the species.

Description
Despite its common name, the Panamanian golden frog is a true toad, a member of the family Bufonidae. It was first described as a subspecies of Atelopus varius, but is now classified as a separate species. The Panamanian golden frog is a national symbol and is considered to be one of the most beautiful frogs in Panama. The skin colour ranges from light yellow-green to bright gold, with some individuals exhibiting black spots on their backs and legs. Females are generally larger than males; females typically range from in length and in weight, with males between in length and in weight. Toxicity The Panamanian golden frog has a variety of toxins, including steroidal bufadienolides and guanidinium alkaloids of the tetrodotoxin class. One of the latter, zetekitoxin AB, has been found to be a blocker of voltage-dependent sodium channels several orders of magnitude more potent than its analog saxitoxin. Their toxin is water-soluble and affects the nerve cells of anyone who comes in contact with it. Panamanian golden frogs use this toxin to protect themselves from most predators. Due to the risk of testing the poison on humans, it has been done with mice. Large doses can be fatal in 20 or 30 minutes. Death is preceded by clonic (grand mal) convulsions until the functions of the circulatory and respiratory systems cease. ==Distribution==
Distribution
The Panamanian golden frog is endemic to Panama, living close to mountain streams on the eastern side of the Tabasará mountain range in the Coclé and Panamá provinces. Individuals are kept in captive-breeding programmes in more than 50 institutions across North America and Panama. ==Ecology==
Ecology
The lifespan of the Panamanian golden frog is 12 years. When encountering another male, male frogs will wave their forefeet as a sign of defense. ==Conservation==
Conservation
The Panamanian golden frog began vanishing from its high mountain forests in the late 1990s, prompting a scientific investigation and rescue process that continues today. It was filmed for the last time in the wild in 2006 by the BBC Natural History Unit for the series Life in Cold Blood by David Attenborough. Additional factors, such as habitat loss and pollution, may have also played a role. If a cold period occurs, the behavior and immunity of the frogs may change around the same time more spores are released. When these frogs are infected with the fungus, their body temperatures rise to fight off the fungus. However, even if the infection leaves the frogs and body temperatures return to previous normal levels, the infection can re-emerge. Another study found that dry conditions added an average 25 days to the lifespan of infected individuals, while higher temperatures only added 4 days. Not only do these frogs face the threat of the fungal disease, but they also are threatened by human development. As trees are cleared for housing and urbanization, the habitat of A. zeteki is destroyed. Other threats include encroachment by agriculture, pollution, pet trade, and aquaculture. Two significant efforts to save these frogs have been made. The Amphibian Recovery Conservation Coalition, which started in 2004, exported the endangered amphibians to the US, believing it was a better environment for the endangered species. In 2005, the Houston Zoo established the El Valle Amphibian Conservation Center (EVACC) in Panama, so the endangered frogs could have protected facilities in their native country. EVACC has become a tourist attraction and the populations of the housed species are watched closely by researchers. In early 2006, EVACC exceeded its capacity for housing golden frogs. In order to continue collection efforts, a partnership was formed with the Hotel Campestre in El Valle whereby rooms 28 and 29 of the resort were loaded with terrariums as a stop-gap measure. This was one of several conservation efforts covered in Elizabeth Kolbert's The Sixth Extinction: An Unnatural History. Over 300 frogs were kept in the so-called "golden frog hotel" and treated to daily cleansing rinses, 24-hour room service, and exotic lunches of specialty crickets until space could be made available in the EVACC. Experiment Prior to the spread of the Batrachochytrium dendrobatidis fungus into Panamanian golden frog habitat, conservation organizations collected Panamanian golden frogs and placed them in captive survival assurance colonies. The skin of amphibians is host to a diverse resident bacterial community, which acts as a defense mechanism in some amphibians to inhibit pathogens. Researchers characterized the bacterial community from wild and captive Panamanian golden frogs originating from the same population with sequencing to assess how long-term captivity has affected this community. Species richness, phylogenetic diversity, and community structure of the skin microbiota were found to be significantly different between wild and captive Panamanian golden frogs. However, after approximately eight years of living in captivity, the offspring of the original captive Panamanian golden frogs still shared 70% of their microbial community with wild frogs. These results demonstrate that host-associated microbial communities can be significantly altered by captive management, but most of the community composition can be preserved. Reintroduction efforts from captive assurance colonies are unlikely to be successful without the development of methods to control chytridiomycosis in the wild. Researchers have experimented by using Janthinobacterium lividum to control the skin temperature in hopes the fungus would be kept at bay. It seemed to be a protective treatment at the early stage, but the frogs began to die after two weeks as the Batrachochytrium dendrobatidis is the causative agent of chytridiomycosis. Other methods of focusing on the phenotypic and genetic concordance to do the conservation have been used. Researchers designated A. varius and A. zeteki as separate species, but they are poorly sorted based on physical characteristics, and better sorted based on mitochondrial DNA. ==In culture==
In culture
The Panamanian golden frog is something of a national symbol, appearing on state lottery tickets and in local mythology. When the toad dies, it is thought to turn to gold and to bring good luck to those fortunate enough to see it. In 2010, the Panamanian government passed legislation recognizing August 14 as "National Golden Frog Day". The main celebration event is marked annually by a parade in the streets of El Valle de Anton, and a display of Panamanian golden frogs at the El Valle Amphibian Conservation Center in the El Nispero Zoo, El Valle. The highly toxic skin of the frog has also been used for centuries by the native people of the Panamanian forests for arrow poison. ==See also==
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