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Pelican

Pelicans are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before swallowing. They have predominantly pale plumage, except for the brown and Peruvian pelicans. The bills, pouches, and bare facial skin of all pelicans become brightly coloured before the breeding season.

Taxonomy and systematics
Etymology The name comes from the Ancient Greek word pelekan (πελεκάν), which is itself derived from the word pelekys (πέλεκυς) meaning "axe". In classical times, the word was applied to both the pelican and the woodpecker. Taxonomy history The genus Pelecanus was first formally described by Carl Linnaeus in his landmark 1758 10th edition of Systema Naturae. He described the distinguishing characteristics as a straight bill hooked at the tip, linear nostrils, a bare face, and fully webbed feet. This early definition included frigatebirds, cormorants, and sulids, as well as pelicans. The family Pelecanidae was later introduced (as Pelicanea) by the French polymath Constantine Samuel Rafinesque in 1815. Order and related taxa Pelicans give their name to the Pelecaniformes, an order which has undergone significant revision. Tropicbirds (now Phaethontiformes), darters, cormorants, gannets, boobies, and frigatebirds (now Suliformes), all traditional members of the order, have since been removed from Pelecaniformes. In their place, herons, ibises, spoonbills, the hamerkop, and the shoebill have now been added into the Pelecaniformes. Together, they form a distinct clade within Pelecaniformes, although their precise evolutionary relationships remain under study. Evolution and fossil record The oldest known pelican fossil is Eopelecanus aegyptiacus, a tibiotarsus from the late Eocene (Priabonian) the Birket Qarun Formation in the Wadi El Hitan in Egypt (~36 million years ago). It shows striking similarities with modern species. Both fossils show a beak nearly morphologically identical to that of present-day pelicans. P. odessanus, Widhalm, 1886 • North America: P. halieus, Wetmore, 1933; P. schreiberi, Olson, 1999 • Asia: P. cautleyi, Davies, 1880; P. sivalensis, Davies, 1880 • Australia: P. cadimurka, Rich & van Tets, 1981; P. tirarensis, Miller, 1966 Controversial and dubious fossil assignments Protopelicanus (Late Eocene) – Once considered a possible early pelecaniform, this bird might instead belong to the Pelagornithidae (pseudotooth birds) or another unrelated aquatic lineage. It is not generally accepted as a member of Pelecanidae. Extant species and phylogeny Species overview There are eight extant species of pelicans, which were historically divided into two groups based on plumage colouration and nesting behavior. One group includes four ground-nesting species with predominantly white plumage—the Australian, Dalmatian, great white, and American white pelicans. The other group consists of four species with grey or brown plumage that nest either in trees or on coastal rocks—the pink-backed, spot-billed, brown, and Peruvian pelicans. However, species with similar plumage and nesting behavior are found in both groups, indicating that these traits do not reflect deep evolutionary divisions. Genetic analyses using mitochondrial and nuclear DNA have revealed a different picture of pelican relationships. These studies support the existence of two major clades: a New World clade, comprising the American white, brown, and Peruvian pelicans, and an Old World clade that includes the Dalmatian, pink-backed, spot-billed, Australian, and great white pelicans. This phylogeny suggests that pelicans evolved in the Old World and later colonized the Americas. Furthermore, it indicates that nesting behavior is more strongly influenced by body size than by genetic lineage. List of living species ==Description==
Description
. The knob on its bill develops before the breeding season. with a chick in a nest in Chesapeake Bay, Maryland, US. This species will nest on the ground when no suitable trees are available. displaying the extent of its throat pouch (Lakes Entrance, Victoria) Pelicans are very large birds with very long bills characterised by a downcurved hook at the end of the upper mandible, and the attachment of a huge gular pouch to the lower. The slender rami of the lower bill and the flexible tongue muscles form the pouch into a basket for catching fish, and sometimes rainwater, They have a long neck and short stout legs with large, fully webbed feet. Although they are among the heaviest of flying birds, they are relatively light for their apparent bulk because of air pockets in the skeleton and beneath the skin, enabling them to float high in the water. Males are generally larger than females and have longer bills. the longest of any bird. The plumage of immature pelicans is darker than that of adults. Newly hatched chicks are naked and pink, darkening to grey or black after four to 14 days, then developing a covering of white or grey down. The air sacs serve to keep the pelican remarkably buoyant in the water and may also cushion the impact of the pelican's body on the water surface when they dive from flight into water to catch fish. Superficial air sacs may also help to round body contours (especially over the abdomen, where surface protuberances may be caused by viscera changing size and position) to enable the overlying feathers to form more effective heat insulation and also to enable feathers to be held in position for good aerodynamics. ==Distribution and habitat==
Distribution and habitat
Modern pelicans are found on all continents except Antarctica. They primarily inhabit warm regions, although breeding ranges extend to latitudes of 45° South (Australian pelicans in Tasmania) and 60° North (American white pelicans in western Canada). Birds of inland and coastal waters, they are absent from polar regions, the deep ocean, oceanic islands (except the Galapagos), and inland South America, as well as from the eastern coast of South America from the mouth of the Amazon River southwards. although their scarcity and isolated occurrence suggests that these remains may have merely been vagrants from Australia (much as is the case today). ==Behaviour and ecology==
Behaviour and ecology
Pelicans swim well with their strong legs and their webbed feet. They rub the backs of their heads on their preen glands to pick up an oily secretion, which they transfer to their plumage to waterproof it. Hence, substantial energy is saved while flying. Adult pelicans rely on visual displays and behaviour to communicate, particularly using their wings and bills. Agonistic behaviour consists of thrusting and snapping at opponents with their bills, or lifting and waving their wings in a threatening manner. Adult pelicans grunt when at the colony, but are generally silent elsewhere or outside breeding season. Conversely, colonies are noisy, as chicks vocalise extensively. In all species, copulation takes place at the nest site; it begins shortly after pairing and continues for three to ten days before egg-laying. The male brings the nesting material, in ground-nesting species (which may not build a nest) sometimes in the pouch, and in tree-nesting species crosswise in the bill. The female then heaps the material up to form a simple structure. Sometimes before, but especially after being fed the pelican chick may seem to "throw a tantrum" by loudly vocalizing and dragging itself around in a circle by one wing and leg, striking its head on the ground or anything nearby and the tantrums sometimes end in what looks like a seizure that results in the chick falling briefly unconscious; the reason is not clearly known, but a common belief is that it is to draw attention to itself and away from any siblings who are waiting to be fed. but occasionally amphibians, turtles, crustaceans, insects, birds, and mammals are also eaten. The size of the preferred prey fish varies depending on pelican species and location. For example, in Africa, the pink-backed pelican generally takes fish ranging in size from fry up to and the great white pelican prefers somewhat larger fish, up to , but in Europe, the latter species has been recorded taking fish up to . Although all pelican species may feed in groups or alone, the Dalmatian, pink-backed, and spot-billed pelicans are the only ones to prefer solitary feeding. When fishing in groups, all pelican species have been known to work together to catch their prey, and Dalmatian pelicans may even cooperate with great cormorants. Pelicans in their turn sometimes snatch prey from other waterbirds. The Australian and American white pelicans may feed by low plunge-dives landing feet-first and then scooping up the prey with the beak, but they—as well as the remaining pelican species—primarily feed while swimming on the water. and "anything from insects and small crustaceans to ducks and small dogs". as well as crowned cormorants, kelp gulls, greater crested terns, and African penguins on Dassen Island and elsewhere. The Australian pelican, which is particularly willing to take a wide range of prey items, has been recorded feeding on young Australian white ibis, and young and adult grey teals and silver gulls. Brown pelicans have been reported preying on young common murres in California and the eggs and nestlings of cattle egrets and nestling great egrets in Baja California, Mexico. Peruvian pelicans in Chile have been recorded feeding on nestlings of imperial shags, juvenile Peruvian diving petrels, and grey gulls. Cannibalism of chicks of their own species is known from the Australian, brown, and Peruvian pelicans. Non-native great white pelicans have been observed swallowing city pigeons in St. James's Park in London, England. ==Status and conservation==
Status and conservation
Populations Globally, pelican populations are adversely affected by these main factors: declining supplies of fish through overfishing or water pollution, destruction of habitat, direct effects of human activity such as disturbance at nesting colonies, hunting and culling, entanglement in fishing lines and hooks, and the presence of pollutants such as DDT and endrin. Most species' populations are more or less stable, although three are classified by the IUCN as being at risk. All species breed readily in zoos, which is potentially useful for conservation management. '', Tortuga Bay, Island of Santa Cruz, Galápagos The combined population of brown and Peruvian pelicans is estimated at 650,000 birds, with around 250,000 in the United States and Caribbean, and 400,000 in Peru. The National Audubon Society estimates the global population of the brown pelican at 300,000. Numbers of brown pelican plummeted in the 1950s and 1960s, largely as a consequence of environmental DDT pollution, and the species was listed as endangered in the US in 1970. With restrictions on DDT use in the US from 1972, its population has recovered, and it was delisted in 2009. The Peruvian pelican is listed as near threatened because, although the population is estimated by BirdLife International to exceed 500,000 mature individuals, and is possibly increasing, it has been much higher in the past. It declined dramatically during the 1998 El Niño event and could experience similar declines in the future. Conservation needs include regular monitoring throughout the range to determine population trends, particularly after El Niño years, restricting human access to important breeding colonies, and assessing interactions with fisheries. The spot-billed pelican has an estimated population between 13,000 and 18,000 and is considered to be near threatened in the IUCN Red List of Threatened Species. Numbers declined substantially during the 20th century, one crucial factor being the eradication of the important Sittaung valley breeding colony in Burma through deforestation and the loss of feeding sites. The chief threats it faces are from habitat loss and human disturbance, but populations have mostly stabilised following increased protection in India and Cambodia. The pink-backed pelican has a large population ranging over much of sub-Saharan Africa. In the absence of substantial threats or evidence of declines across its range, its conservation status is assessed as being of least concern. Regional threats include the drainage of wetlands and increasing disturbance in southern Africa. The species is susceptible to bioaccumulation of toxins and the destruction of nesting trees by logging. The American white pelican has increased in numbers, However, whether its numbers have been affected by exposure to pesticides is unclear, as it has also lost habitat through wetland drainage and competition with recreational use of lakes and rivers. The Dalmatian pelican has a population estimated at between 10,000 and 20,000 following massive declines in the 19th and 20th centuries. The main ongoing threats include hunting, especially in eastern Asia, disturbance, coastal development, collision with overhead power lines, and the over-exploitation of fish stocks. It is listed as near threatened by the IUCN Red List of Threatened Species as the population trend is downwards, especially in Mongolia, where it is nearly extinct. However, several European colonies are increasing in size and the largest colony for the species, at the Small Prespa Lake in Greece, has reached about 1,400 breeding pairs following conservation measures. Widespread across Australia, Overall population numbers fluctuate widely and erratically depending on wetland conditions and breeding success across the continent. The species is assessed as being of least concern. Culling and disturbance Pelicans have been persecuted by humans for their perceived competition for fish, despite the fact that their diet overlaps little with fish caught by people. Great white pelicans on Dyer Island, in the Western Cape region of South Africa, were culled during the 19th century because their predation of the eggs and chicks of guano-producing seabirds was seen to threaten the livelihood of the guano collectors. Poisoning and pollution DDT pollution in the environment was a major cause of decline of brown pelican populations in North America in the 1950s and 1960s. It entered the oceanic food web, contaminating and accumulating in several species, including one of the pelican's primary food fish – the northern anchovy. Its metabolite DDE is a reproductive toxicant in pelicans and many other birds, causing eggshell thinning and weakening, and consequent breeding failure through the eggs being accidentally crushed by brooding birds. Since an effective ban on the use of DDT was implemented in the US in 1972, the eggshells of breeding brown pelicans there have thickened and their populations have largely recovered. In the late 1960s, following the major decline in brown pelican numbers in Louisiana from DDT poisoning, 500 pelicans were imported from Florida to augment and re-establish the population; over 300 subsequently died in April and May 1975 from poisoning by the pesticide endrin. About 14,000 pelicans, including 7,500 American white pelicans, perished from botulism after eating fish from the Salton Sea in 1990. As waterbirds that feed on fish, pelicans are highly susceptible to oil spills, both directly by being oiled and by the impact on their food resources. A 2007 report to the California Fish and Game Commission estimated that during the previous 20 years, some 500–1,000 brown pelicans had been affected by oil spills in California. Where pelicans interact with fishers, through either sharing the same waters or scavenging for fishing refuse, they are especially vulnerable to being hooked and entangled in both active and discarded fishing lines. Fish hooks are swallowed or catch in the skin of the pouch or webbed feet, and strong monofilament fishing line can become wound around bill, wings, or legs, resulting in crippling, starvation, and often death. Local rescue organisations have been established in North America and Australia by volunteers to treat and rehabilitate injured pelicans and other wildlife. Parasites and disease As with other bird families, pelicans are susceptible to a variety of parasites. Avian malaria is carried by the mosquito Culex pipiens, and high densities of these biting insects may force pelican colonies to be abandoned. Leeches may attach to the vent or sometimes the inside of the pouch. A study of the parasites of the American white pelican found 75 different species, including tapeworms, flukes, flies, fleas, ticks, and nematodes. The brown pelican has a similarly extensive range of parasites. The nematodes Contracaecum multipapillatum and C. mexicanum and the trematode Ribeiroia ondatrae have caused illness and mortality in the Puerto Rican population, possibly endangering the pelican on this island. Many pelican parasites are found in other bird groups, but several lice are very host-specific. In May 2012, hundreds of Peruvian pelicans were reported to have perished in Peru from a combination of starvation and roundworm infestation. ==Symbolism and cultural significance==
Symbolism and cultural significance
The pelican has played a prominent symbolic role in human cultures across time and geography. From ancient Egypt to Indigenous Australia, and from Christian allegory to modern logos and mascots, pelicans have been interpreted as emblems of protection, sacrifice, care, and transformation. In Jewish dietary law, pelican is not considered kosher (fit for consumption), as it is a type of seabird and therefore considered an unclean animal. An origin myth from the Murri people of Queensland, cited by Andrew Lang, describes how the Australian pelican acquired its black and white plumage. The story tells that the pelican was once a black bird. During a flood, he made a canoe to save drowning people. He fell in love with a woman and decided to save her, but she and her friends tricked him and escaped. The pelican consequently began preparing to go to war against them by daubing himself with white clay as war paint. Before he had finished, another pelican, on seeing such a strange piebald creature, killed him with its beak, and all such pelicans have been black and white ever since. The Moche people of ancient Peru worshipped nature. They placed emphasis on animals and often depicted pelicans in their art. Christian symbolism poster Myth of self-sacrifice The Physiologus, a didactic Christian text from the 3rd or 4th century, claims that pelicans kill their young when they grow and strike their parents in the face, but then the mother laments them for three days, after which she strikes her side and brings them back to life with her blood. The Physiologus explains this as mirroring the pain inflicted on God by people's idolatry, and the self-sacrifice of Jesus on the cross which redeems the sinful (see the blood and water gushing from the wound in his side). supplementing the image of the lamb and the flag. This mythical characteristic is referenced in the hymn "Adoro te devote" ("Humbly We Adore Thee"), where in the penultimate verse, Saint Thomas Aquinas describes Christ as the loving divine pelican, one drop of whose blood can save the world. Similarly, the 1678 Christian allegorical novel ''The Pilgrim's Progress'' describes how "the pelican pierce[s] her own breast with her bill … to nourish her young ones with her blood, and thereby to show that Christ the blessed so loveth his young, his people, as to save them from death by his blood." The pelican is featured in many Christian artworks, especially in Europe. For example, the first (1611) edition of the King James Bible contains a depiction of a pelican feeding her young in an oval panel at the bottom of the title page. (circa 1573), in which Elizabeth I wears the medieval symbol of the pelican on her chest Elizabeth I and the Church Elizabeth I of England adopted the symbol, portraying herself as the "mother of the Church of England". A portrait of her called the Pelican Portrait was painted around 1573, probably by Nicholas Hilliard. Heraldry and symbolism depict a "pelican in her piety" both in the crest and shield.|left Heraldic imagery Pelicans have featured extensively in heraldry, generally using the Christian symbolism of the pelican as a caring and self-sacrificing parent. Heraldic images featuring a "pelican vulning" refers to a pelican injuring herself, while a "pelican in her piety" refers to a female pelican feeding her young with her own blood. The King of Portugal John II adopted the pelican as is own personal sygil while he was Infante, evoking the Christian symbology to equate the sacrifice of his blood to feed the nation. The pelican as a symbol also became synonymous with the increasing charity efforts of the Santas Casas da Misericórdia during his reign and the reconstruction of the Hospital das Caldas da Rainha and the Hospital Real de Todos-os-Santos, which were mainly patronaged by his wife D. Leonor. Public symbols The heraldic pelican also ended up as a pub name and image, though sometimes with the image of the ship Golden Hind. Sir Francis Drake's famous ship was initially called Pelican, and adorned the British halfpenny coin. Emblems and logos in institutions Educational institutions Pelicans are widely used as emblems by educational institutions, especially universities. In Louisiana, the bird adorns the seals of Louisiana State University, Tulane University, Louisiana Tech University, the University of Louisiana at Lafayette, Loyola University New Orleans, Southeastern Louisiana University, and Southern University. The seal of the Packer Collegiate Institute, depicting a pelican feeding her young, has been in use since 1885. The medical faculties of Charles University in Prague also have a pelican as their emblem, invoking the bird's long-standing association with self-sacrifice in Christian symbolism. The image became also linked to the medieval religious feast of Corpus Christi. The universities of Oxford and Cambridge each have colleges named for the religious festival nearest the dates of their establishment, and Corpus Christi College, Oxford, feature pelicans on their coats of arms. Sports teams In sports, the pelican serves as a mascot and logo for various teams and university athletics. It is the mascot of the New Orleans Pelicans NBA team, the Lahti Pelicans ice hockey team, Tulane University, and the University of the West Indies. Commercial and nonprofit organizations company, Pelikan. Inspired by the coat of arms of its former owner Günther Wagner. The name and image were also employed by Pelican Books, an imprint of nonfiction titles published by Penguin Books. Similarly, Sanquin, the nonprofit organization responsible for blood supply in the Netherlands, uses a stylized pelican in its logo, continuing this humanitarian association.|left coin.|left National and regional symbols As a cultural symbol of nations and states, the pelican holds prominent status. The great white pelican is the national bird of Romania. The brown pelican is the national bird of three Caribbean countries—Saint Kitts and Nevis, Barbados, and Sint Maarten—and features on their coats of arms. A Dalmatian pelican is also depicted on the reverse of the Albanian 1 lek coin, issued in 1996. In the United States, it is the state bird of Louisiana, which is colloquially known as the Pelican State; the bird appears on both the state flag and state seal. '', a member of Archaeidae, also known as pelican spiders. A lateral view of a female and its genitalia. Namesakes in nature Archaeidae, a family of spiders, are known as pelican spiders. The original version ran: ==Notes==
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