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Passalidae

Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles", "betsy bugs", or "horned passalus beetles". Nearly all of the approximately 500 species are tropical; species found in North America are notable for their size, ranging from 20 to 43 mm, for having a single "horn" on the head, and for a form of social behavior unusual among beetles.

Genera
– – Aponelides – – – – CacoiusCeracupes – – Cicernonius – – CylindrocaulusDidimusEpishenusEpisphenoidesEumelosomusFlaminius – – HeliscusHincksius – – – – MalagasulusMastochilusNeleidesNeleuopsNeluesNinusOdontotaeniusOgygesOileusOphrygoniusOrgyesParatiberioidesPassalusPassipassalus – – – PelopsPentalobus – – PetrejusPharochilusPhoroneusPlesthenusPleurariusPopiliusProculejus – – Protomocoelus – – – Ptichopus – – Rhodocanthopus – – – – Spurius – – – TiberioidesTrichopleurusUnduliferVerres – – Vindex – – ==Selected species==
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