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Porcupine

Porcupines are large rodents with coats of sharp spines, or quills, that protect them against predators. The term covers two families of animals, the Old World porcupines in the family Hystricidae, and the New World porcupines in the family Erethizontidae. Both families display superficially similar coats of rigid or semirigid quills, which are modified hairs composed of keratin, and belong to the infraorder Hystricognathi within the diverse order Rodentia. The two groups are distinct and are not closely related to each other within the Hystricognathi. The largest species of porcupine is the third-largest living rodent in the world, after the capybara and beaver.

Terminology and etymology
The word "porcupine" comes from the Latin + , from Old Italian porcospino, . A regional American name for the animal is "quill-pig". A baby porcupine is known as a porcupette. When born, a porcupette's quills are soft hair; they harden within a few days, forming the sharp quills of adults. ==Taxonomy and evolution==
Taxonomy and evolution
Classification A porcupine is any of 29 species of rodents belonging to the families Erethizontidae (genera: Coendou, Erethizon, and Chaetomys) or Hystricidae (genera: Atherurus, Hystrix, and Trichys). The two families of porcupines are quite different, and although both belong to the Hystricognathi branch of the vast order Rodentia, they are not closely related. Porcupines are distributed into two evolutionarily independent groups within the infraorder Hystricognathi, part of the suborder Hystricomorpha of the Rodentia: • Parvorder Phiomorpha • Family Hystricidae: Old World porcupines • African brush-tailed porcupine, Atherurus africanusAfrican crested porcupine, Hystrix cristataAsiatic brush-tailed porcupine, Atherurus macrourusCape porcupine, Hystrix africaeaustralisIndian crested porcupine, Hystrix indicusMalayan porcupine, Hystrix brachyuraSunda porcupine, Hystrix javanicaSumatran porcupine, Hystrix (Thecurus) sumatraeThick-spined porcupine, Hystrix (Thecurus) crassispinisPhilippine porcupine, Hystrix (Thecurus) pumilisLong-tailed porcupine, Trichys fasciculata • Parvorder Caviomorpha • Family Erethizontidae: New World porcupines • North American porcupine, Erethizon dorsatumBrazilian porcupine, Coendou prehensilisBicolored-spined porcupine, Coendou bicolorAndean porcupine, Coendou quichuaBlack dwarf (Koopman's) porcupine, Coendou nycthemeraRothschild's porcupine, Coendou rothschildiSanta Marta porcupine, Coendou sanctemartaeMexican hairy dwarf porcupine, Coendou mexicanusParaguaian hairy dwarf porcupine, Coendou spinosusBahia porcupine, Coendou insidiosusBrown hairy dwarf porcupine, Coendou vestitusStreaked dwarf porcupine, Coendou ichillusBlack-tailed hairy dwarf porcupine, Coendou melanurusRoosmalen's dwarf porcupine, Coendou roosmalenorumFrosted hairy dwarf porcupine, Coendou pruinosusStump-tailed porcupine, Coendou rufescensBristle-spined porcupine, Chaetomys subspinosus (sometimes considered an echimyid) Evolution Studies on the physical characteristics of New World porcupines initially described them as being the most basal (earliest to diverge) group among the hystricognaths, Dorothée Huchon and Emmanuel Douzery wrote in 2001 on the diversification of the hystricognaths, reporting that molecular phylogenetic studies using nuclear genes (e.g., the von Willebrand factor gene) suggested that the Hystricidae (Old World porcupines), Phiomorpha, and Caviomorpha (which includes New World porcupines) lineages split some 63 to 43 million years ago. This supported a single colonization event of South America by hystricognaths. Fossils belonging to the genus Hystrix date back to the late Miocene of Africa and Asia. One of the oldest known porcupine fossils is Hystrix primigenia, an Old World porcupine that lived in the late Miocene to the Pliocene; the oldest known porcupine fossil in Asia, Hystrix lufengensis, dates back , also in the late Miocene. == Biology ==
Biology
, the Cape porcupine Old World compared with New World species The 11 Old World porcupines tend to be fairly large and have spines grouped in clusters. The two subfamilies of New World porcupines are mostly smaller (although the North American porcupine reaches about in length and ), have their quills attached singly rather than grouped in clusters, and are excellent climbers, spending much of their time in trees. The New World porcupines evolved their spines independently (through convergent evolution) and are more closely related to several other families of rodents than they are to the Old World porcupines. Description Porcupines are rodents of varying fur color that are characterized by their abundance of protruding spines, or quills, found all along the head and body of the animal. Some species have quills that extend from the tail, as well. They vary in size considerably: an adult Andean porcupine weighs roughly , while the crested porcupine can grow to weigh up to . Longevity Porcupines have relatively long lifespans and hold the record for being the longest-living rodent, with one individual named Cooper living over 32 years. Diet The North American porcupine is an herbivore and often climbs trees for food; it eats leaves, herbs, twigs, and green plants such as clover. In the winter, it may eat bark. The African porcupine is not a climber; instead, it forages on the ground. but sometimes forages for food during the day, eating bark, roots, fruits, berries, and farm crops. Porcupines have become a pest in Kenya and are eaten as a delicacy. Defense Defensive behaviour displays in a porcupine depend on sight, scent, and sound. Often, these displays are shown when a porcupine becomes agitated or annoyed. The four main displays seen in a porcupine are (in order from least to most aggressive) quill erection, teeth clattering, odour emission, and attack. A porcupine's colouring aids in part of its defence, as most of the predators are nocturnal and colour-blind. A porcupine's markings are black and white. The dark body and coarse hair of the porcupine are dark brown/black and when quills are raised, present a white strip down its back mimicking the look of a skunk. This, along with the raising of the sharp quills, deters predators. Along with the raising of the quills, porcupines clatter their teeth to warn predators not to approach. The incisors vibrate against each other, the strike zone shifts back, and the cheek teeth clatter. This behaviour is often paired with body shivering, which is used to further display the dangerous quills. The use of odour is when the sight and sound have failed. An unpleasant scent is produced from the skin above the tail in times of stress and is often seen with a quill erection. If these processes fail, the porcupine attacks by running sideways or backwards into predators. A porcupine's tail can also be swung in the direction of the predator; if contact is made, the quills could be impaled into the predator, causing injury or death. Quills Porcupines' quills, or spines, take on various forms depending on the species, but all are modified hairs coated with thick plates of keratin and are embedded in the skin musculature. Old World porcupines have quills embedded in clusters, whereas in New World porcupines, single quills are interspersed with bristles, underfur, and hair. Quills are released by contact or may drop out when the porcupine shakes its body. New quills grow to replace lost ones. Some possible antibiotic properties are within the quills, specifically associated with the free fatty acids coating them. The antibiotic properties are believed to aid a porcupine that has suffered from self-injury. ==Ecology==
Ecology
Behavior Porcupines are slow-moving animals. All are social to some degree, though only Old World porcupines are known to form clans or family units. Habitat and range Porcupines occupy a small range of habitats in tropical and temperate parts of Asia, Southern Europe, Africa, and North and South America. They live in forests and deserts, rocky outcrops, and hillsides. Some New World porcupines live in trees, but Old World porcupines prefer a rocky environment. Porcupines can be found on rocky areas up to high. They are generally nocturnal, but are occasionally active during daylight. ==Relationships with humans==
Relationships with humans
Use as food and clothing displayed at the Museo de Arte Popular in Mexico City Porcupines are seldom eaten in Western culture, but are eaten often in Southeast Asia, particularly Vietnam, where the prominent use of them as a food source has contributed to declines in their populations.