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American bullfrog

The American bullfrog, often simply known as the bullfrog in Canada and the United States, is a large true frog native to eastern North America. It typically inhabits large permanent water bodies such as swamps, ponds, and lakes. Bullfrogs can also be found in manmade habitats such as pools, koi ponds, canals, ditches, reservoirs and culverts. The bullfrog gets its name from the sound the male makes during the breeding season, which sounds similar to a bull bellowing. They are generalist carnivores, consuming a wide variety of prey. In turn, they are eaten by predators such as alligators, snakes, and birds.

Taxonomy
The specific name, catesbeiana (feminine) or catesbeianus (masculine), is in honor of English naturalist Mark Catesby. Some authorities use the scientific name, Lithobates catesbeiana, although others prefer Rana catesbeiana. Cladogram of the position of the American bullfrog within Lithobates, after Martínez‐Gil et al. 2025:{{clade| ==Description==
Description
The dorsal (upper) surface of the bullfrog has an olive-green background color, either plain or with mottling and banding of grayish brown. The ventral (under) surface is off-white blotched with yellow or gray. Often, a marked contrast in color is seen between the green upper lip and the pale lower lip. Bullfrogs are sexually dimorphic, with males being smaller than females and having yellow throats. Males have tympana larger than their eyes, whereas the tympana in females are about the same size as the eyes. Also, such mechanism relieves the tongue's musculature from physiological constraints such as limited peak power output - mechanical efficiency and thermal dependence by uncoupling the activation of the depressor mandibulae's contractile units from actual muscular movement. In other words, the kinematic parameters developed by contribution of the elastic structures differ from those developed by muscular projection, accounting for the difference in velocity, power output, and thermal dependence. ==Distribution==
Distribution
The bullfrog is originally native to eastern North America, where it is commonly found in every U.S. state east of the Mississippi River. Its natural range extends from the eastern Canadian Maritime Provinces to as far west as Idaho and Texas, and as far north as Michigan (including the Upper Peninsula), Minnesota and Montana; it is largely absent in North Dakota. Within North America, the bullfrog has been introduced to Nantucket island, as well as portions of the western U.S., including Arizona, California, Colorado, Idaho, Nevada, New Mexico, Oregon, Utah, Washington and Wyoming. In these states, it is considered to be an invasive species, as concerns exist that it may outcompete or prey upon native species of reptiles and amphibians, disrupting the delicate ecological balance of certain areas. Bullfrogs have been found to feed on the young of several snakes, including the California endemic giant garter snake, a threatened species. In early 2023, the Utah Department of Natural Resources began tweeting tips on how to catch and cook bullfrogs in an effort to encourage residents to help control the growing population by catching the invasive frogs for food. The bullfrog has also been introduced to Hawaii, South America, Asia, the Caribbean, and Europe for various purposes including frog farming and population control of other species. Specifically, the bullfrog has been introduced to the extreme south of British Columbia, Canada, nearly every state in Mexico, as well as Belgium, Cuba, the Dominican Republic, Haiti, Italy, Jamaica, the Netherlands, and Puerto Rico. It is also found in Argentina, Brazil, China, Colombia, Japan, South Korea, Uruguay and Venezuela. The reasons for introducing the bullfrog to these areas have largely been intentional, either to provide humans with a source of food or as biological control agents. In addition, the unintended escape of frogs from breeding establishments or scientific research facilities, captive escapees or released pets are also a possibility. ==Biology==
Biology
Ecology Bullfrogs are voracious, opportunistic, ambush predators that prey on any small animal they can overpower and consume. Bullfrog stomachs have been found to contain rodents, small lizards and snakes, other frogs and toads, other amphibians, crayfish, other crustaceans, small birds, scorpions, tarantulas and bats, as well as the many types of invertebrates, such as snails, worms and insects, which are the usual food of ranid frogs. These studies revealed the bullfrog's diet to be unique among North American ranids in the inclusion of a large percentage of aquatic animals, such as fish, tadpoles, ram's horn snails, and dytiscid beetles, as well as the aquatic eggs of fish, frogs, insects, or salamanders. Bullfrogs are able to capture large, strong prey because of the powerful grip of their jaws after the initial ranid tongue strike. However, there is a correlation found with size of prey relative to body size of the bullfrog. Juveniles and adults typically go after prey that is relative to their own body size. The bullfrog is able to make allowance for light refraction at the water-air interface by striking at a position posterior to the target's perceived location. The comparative ability of bullfrogs to capture submerged prey, compared to that of the green frog, leopard frog, and wood frog (L. clamitans, L. pipiens, and L. sylvaticus, respectively) was also demonstrated in laboratory experiments. The eggs and larvae are unpalatable to many salamanders and fish, but the high levels of activity of the tadpoles may make them more noticeable to a predator not deterred by their unpleasant taste. Humans hunt bullfrogs as game and consume their legs. Adult frogs try to escape by splashing and leaping into deep water. A trapped individual may squawk or emit a piercing scream, which may surprise the attacker sufficiently for the frog to escape. An attack on one bullfrog is likely to alert others in the vicinity to danger and they will all retreat into the safety of deeper water. Bullfrogs may be at least partially resistant to the venom of copperhead (Agkistrodon contortrix) and cottonmouth (Agkistrodon piscivorus) snakes, though these species are known natural predators of bullfrogs as are northern water snakes (Nerodia sipedon). As an invasive species Multiple traits of L. catesbeianus contribute to its competitive ability, especially as an invasive. The generalist diet of the American bullfrog allows for it to consume food in different environments. They directly predate indigenous frog species, leading to numerous frog declines. When observing the contents of American bullfrog stomachs, it was discovered that adult bullfrogs regularly consume predators of bullfrog young, including dragonfly nymphs, garter snakes, and giant water bugs. Thus, the ecological check on American bullfrog juveniles in invaded areas become less effective. L. catesbeianus seems to exhibit traits of immunity or resistance against the antipredator defenses of other organisms, with analysis of stomach contents from New Mexican bullfrogs showing the regular consumption of wasps, with no conditioned avoidance due to the wasps' stingers. Along the Colorado river, L. catesbeianus stomach contents indicate the ability to withstand the discomforting spines of the stickleback fish. Reports of American bullfrogs eating scorpions and rattlesnakes also exist. Analysis of the American bullfrog's realized niche at various sites in Mexico, and comparisons with the niches of endemic frogs show that it is possible that the American bullfrog capable of niche shift, and pose a threat to many endemic Mexican frog species, even those that are not currently in competition with the American bullfrog. Self-sustaining populations of American bullfrogs became established in the United Kingdom around 1999, where their introduction was likely due to accidental escapes and deliberate releases from captivity. These populations appear to be quite small, and are undergoing control by Natural England as the species poses a threat to native amphibians. The American bullfrog has been known to spread the amphibian pathogen Batrachochytrium dendrobatidis among populations that it has been introduced to. Breeding season The bullfrog breeding season typically lasts two to three months. Conversely, females have brief periods of sexual receptivity during the season. In one study, female sexual activity typically lasted for a single night and mating did not occur unless the females initiated the physical contact. The significance of forelimb sexual dimorphism allow males to remain in amplexus with the female for longer durations increasing their chance at reproductive success in the highly competitive mating environment. Females can deposit 1,000 - 40,000 eggs which hatch in 3 - 5 days. These male and female behaviors cause male-male competition to be high within the bullfrog population and sexual selection for the females to be an intense process. This may reflect a lesser number of predators in these locations. As they grow, they tend to move into deeper water. The tadpoles initially have three pairs of external gills and several rows of labial teeth. They pump water through their gills by movements of the floor of their mouths, trapping bacteria, single-celled algae, protozoans, pollen grains, and other small particles on mucus in a filtration organ in their pharynges. As they grow, they begin to ingest larger particles and use their teeth for rasping. They have downward-facing mouths, deep bodies, and tails with broad dorsal and ventral fins. Time to metamorphosis ranges from a few months in the southern part of the range to 3 years in the north, where the colder water slows development. Maximum lifespan in the wild is estimated to be 8 to 10 years, but one frog lived for almost 16 years in captivity. ==Relation to humans==
Relation to humans
The American bullfrog provides a food source, especially in the Southern and some areas of the Midwestern United States. The traditional way of hunting them is to paddle or pole silently by canoe or flatboat in ponds or swamps at night; when the frog's call is heard, a light is shone at the frog which temporarily inhibits its movement. The frog will not jump into deeper water as long as it is approached slowly and steadily. When close enough, the frog is gigged with a multiple-tined spear and brought into the boat. Bullfrogs can also be stalked on land, by again taking great care not to startle them. In some states, breaking the skin while catching them is illegal, and either grasping gigs or hand captures are used. Like most frogs, the hind legs of the bullfrog are the only parts generally eaten. When cooked, they resemble small chicken drumsticks, have a similar flavor and texture and can be prepared in similar ways. During the Great Depression, businessman Dr. Albert Broel promoted bullfrog farming as a way for people experiencing difficult economic times to earn extra money by raising and selling bullfrogs. He published several instructional manuals, including "Frog Raising for Pleasure and Profit," as well as in-person training services. Broel offered to purchase all the adult frogs independent breeders could successfully raise to adult weight. The scheme was, ultimately, unsuccessful for many of the reasons described above. However, the business model of outsourcing livestock raising to independent farmers continues today, especially in the poultry industry. The American bullfrog is used as a specimen for dissection in many biology and anatomy classes in schools across the world. It is the state amphibian of Missouri, Ohio, and Oklahoma. In areas where the American bullfrog is introduced, the population can be controlled by various means. One project (3n-Bullfrog project) uses sterile triploïd (3n) bullfrogs. In Europe, the American bullfrog is included since 2016 in the list of Invasive Alien Species of Union concern (the Union list). This means that this species cannot under any circumstances be imported, bred, transported, commercialized, or intentionally released into the environment in the whole of the European Union. ==References==
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