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Hystricomorpha

Hystricomorpha is a term referring to families and orders of rodents which has had many definitions throughout its history. In the broadest sense, it refers to any rodent with a hystricomorphous zygomasseteric system. This includes the Hystricognathi, Ctenodactylidae, Anomaluridae, and Pedetidae. Molecular and morphological results suggest the inclusion of the Anomaluridae and Pedetidae in Hystricomorpha may be suspect. Based on Carleton & Musser 2005, these two families are discussed here as representing a distinct suborder Anomaluromorpha.

Classification
showing the enlarged infraorbital canal present in most members of the Hystricomorpha: This condition is termed hystricomorphy. The modern definition of Hystricomorpha, also known as Entodacrya or Ctenohystrica, is a taxonomic hypothesis uniting the gundis with the hystricognath rodents. Considerable morphological and strong molecular support exists for this relationship. If true, this hypothesis renders the traditional view of Sciurognathi invalid, as it becomes a paraphyletic group. The hystricomorph rodents, or at least members of Caviomorpha, are sometimes not regarded as rodents. Most molecular and genetic research, however, confirms the monophyly of rodents. Support for rodent polyphyly appears to be a product of long branch attraction. Hystricomorph rodents appeared in South America in the Eocene, a continent which previously had metatherians, xenarthrans, and meridiungulates as the only resident nonflying mammals. They apparently arrived by rafting across the Atlantic from Africa. The same type of migration may have occurred with primates, which also appeared in South America in the Eocene when it was an isolated continent, long before the Great American Interchange. All of this is still controversial, and new scientific discoveries on this subject are published regularly. ==Families==
Families
The following list of families is based on the taxonomy of and , who subjected a number of early fossil rodents to parsimony analysis and recovered support for the Hystricomorpha or Entodacrya hypothesis. Their results rendered the suborder Sciuravida as defined by to be polyphyletic and invalid. The symbol "†" is used to indicate extinct groups. • Suborder Hystricomorpha • †Pseudoneoreomys? • Superfamily Ctenodactyloidea • †ChapattimyidaeCtenodactylidae – gundis • DiatomyidaeLaotian rock rat • †Tamquammyidae • †Yuomyidae • Hystricognathiformes • †TsaganomyidaeHystricognathi – true hystricognaths • †BaluchimyinaePhiomorphaBathyergidae – blesmols • †Bathyergoididae • †DiamantomyidaeHeterocephalidae – naked mole-rats • Hystricidae – Old World porcupines • †Kenyamyidae • †MyophiomyidaePetromuridae – dassie rat • †PhiomyidaeThryonomyidae – cane rats • Caviomorpha – New World hystricognaths • Superfamily CavioideaCaviidae – cavies, capybaras, and guinea pigs • †CephalomyidaeCuniculidae – pacas • Dasyproctidae – agoutis and acouchis • Dinomyidae – pacarana • †Eocardiidae • †Neoepiblemidae • Superfamily ChinchilloideaAbrocomidae – chinchilla rats • Chinchillidae – chinchillas and viscachas • Superfamily ErethizontoideaErethizontidae – New World porcupines • Superfamily OctodontoideaCapromyidae – hutias • Ctenomyidae – tuco-tucos • Echimyidae – spiny rats • †Heptaxodontidae – giant hutias • Myocastoridae – nutria • Octodontidae – degus and relatives == Citations ==
General references
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