Wrottesley was nominated for Parliament for the
general election of 22 March 1768 by Earl Gower as member for
Newcastle-under-Lyme. The borough was regarded as belonging to the Gower Leveson family, who owned much of the property and allowed the tenants to get into serious arrears, as well as providing lavish hospitality during elections. However, the borough had a large electorate and had recently been made less secure by the emergence of a malcontent group of electors, who invited
Robert Clive to challenge the controlling interest. The challenge was unsuccessful and Wrottesley was returned unopposed, alongside
Alexander Forrester, a major member of the
Bedfordite faction of conservative Whigs, placed in the seat at the request of the
Duke John Russell himself. The Gowers were now allied with the Duke of Bedford's faction and this was to determine most of Wrottesley's parliamentary career. Only a month after the election, a vacancy occurred in
Staffordshire, when one member,
George Harry Grey, succeeded his father as
Earl of Stamford. Grey had been a Gower nominee in this
county constituency, where, by informal agreement, the Gowers nominated one member, while leaving the other seat for one of the local
landed gentry. Wrottesley resigned as Member a couple of months later and was assigned to the vacancy by Gower, becoming MP for Staffordshire on 5 July 1768. He represented the county for the remainder of his life, always being returned unopposed. At first Wrottesley almost invariably voted with the government. Such consistency could be achieved only by a political transformation, as the administration itself changed in 1770. For his first two years in Parliament, Wrottesley, the Gowers and the rest of the Bedfordites were involved with the Whig administration of Wrottesley's own brother-in-law, Grafton, a friend and supporter of
William Pitt, 1st Earl of Chatham. The Chathamites favoured an aggressive, anti-French colonial policy, and Wrottesley had made his military career in pursuit of their objectives. Regarding the tensions in North America, however, the Chathamites tended to sympathise with the colonists. The Bedfordite Whigs increasingly adopted the opposite opinion. During the Tory administration of
Frederick North, Lord North, from 1770 onwards, the Bedfordites were allied with the Tories. Wrottesley and the rest of the Gower clients thus changed sides, supporting a hard line against the Americans that led to the
American Revolutionary War. Gower assumed control of the faction when Bedford died in 1771. Wrottesley was still a member of the Army and in 1770 had been promoted to lieutenant colonel. When hostilities began in 1775, he rejoined his regiment, retaining his membership in Parliament. Wrottesley served for three years in the war, returning to England in 1778. By this time, France had formally intervened in the war on the side of the Americans. Wrottesley made his first parliamentary speech after his return on 26 November 1778, during the debate on the Address, which followed the
Speech from the throne at the opening of Parliament. :"(Wrottesley) asked if the House was called upon for unanimity against France? If it was for a war with America, he could not give his approbation to it. All that could be done, he said, had been done. If 50,000 Russians were sent, they could do nothing. He thought New York, Rhode Island, and Halifax should be garrisoned, and the rest of the army brought away." With a speech in December he elaborated his opinion, saying that the British forces were distributed too widely and the lines of communication too stretched for any offensive action. He was not for immediate withdrawal but it was too late for any offensive action. For a year he went no further in his criticisms and supported the government in most votes. However, in a speech on 6 December 1779 he was much more critical. Claiming he was no party man, he nevertheless told ministers: :"A change of measures was now become absolutely necessary. America was lost by their incapacity and misconduct, their obstinacy and their blindness." Wrottesley's evolution followed that of the Bedfordites in general. At the end of 1779, Gower ended the faction's alliance with North, making the end of the government and the war likely. However, Wrottesley continued to support the government in general, voting against opposition attempts to control the expenses of the royal household, but he criticised the government's policy in Ireland, warning that it risked making the same mistakes as in America. On 12 December 1781 Wrottesley voted against a motion from
Sir James Lowther, 5th Baronet to end the war, declaring himself against the total withdrawal implied by the motion. However, on 22 Feb. 1782
Henry Seymour Conway, a veteran of both the
War of the Austrian Succession and
Seven Years' War made a motion against the war. Wrottesley voted for it but it was defeated by the smallest possible margin: 194 votes to 193. Five days later Conway moved a similar motion, again with Wrottesley's endorsement, which was approved by 234 to 215. On 15 March Wrottesley also voted for the motion of no confidence in North's administration made by
Sir John Rous, which finally ended the government. The end of North's government resulted in a government dominated by the
Rockingham Whigs, who had consistently opposed the
American Revolutionary War. However, the unexpected death of
Rockingham in July left
Lord Shelburne trying to hold together an unsteady coalition to bring the war to an end. Wrottesley voted for Shelburne's preliminary efforts for peace on 18 February 1783 – part of a rallying of independent members that allowed negotiations to proceed. Thereafter, Wrottesley abstained from voting on major measures proposed by the
Fox-North Coalition that superseded Shelburne's ministry. He gave general support to
William Pitt the Younger when he became Prime Minister at the end of 1783. Wrottesley's military career progressed just as well while he was in the House of Commons as when he was on active duty. He was promoted to colonel in 1779 and major-general in 1782, and was appointed colonel of the
45th Foot in 1784. He died in 1787, at age 42. ==Family==