) to be phonated through the
vocal folds of the larynx (
phonation) and resonated in the vocal cavities shaped by the
jaw,
soft palate,
lips,
tongue and other articulators (
articulation).
Respiration Respiration is the physical process of gas exchange between an organism and its environment involving four steps (
ventilation, distribution,
perfusion and diffusion) and two processes (inspiration and expiration).
Respiration can be described as the mechanical process of air flowing into and out of the
lungs on the principle of
Boyle's law, stating that, as the volume of a container increases, the air pressure will decrease. This relatively negative pressure will cause air to enter the container until the pressure is equalized. During
inspiration of air, the diaphragm contracts and the lungs expand drawn by
pleurae through surface tension and negative pressure. When the lungs expand, air pressure becomes negative compared to atmospheric pressure and air will flow from the area of higher pressure to fill the lungs. Forced inspiration for speech uses accessory muscles to elevate the
rib cage and enlarge the
thoracic cavity in the vertical and lateral dimensions. During forced expiration for speech, muscles of the
trunk and
abdomen reduce the size of the thoracic cavity by compressing the abdomen or pulling the rib cage down forcing air out of the lungs.
Phonation Phonation is the production of a periodic sound wave by vibration of the
vocal folds. Airflow from the lungs, as well as
laryngeal muscle contraction, causes movement of the vocal folds. It is the properties of tension and elasticity that allow the vocal folds to be stretched, bunched, brought together and separated. During prephonation, the vocal folds move from the abducted to
adducted position.
Subglottal pressure builds and air flow forces the folds apart, inferiorly to superiorly. If the volume of airflow is constant, the velocity of the flow will increase at the area of constriction and cause a decrease in pressure below once distributed. This negative pressure will pull the initially blow open folds back together again. The cycle repeats until the vocal folds are abducted to inhibit phonation or to take a breath.
Articulation In a third process of speech production, articulation, mobile and immobile structures of the face (articulators) adjust the shape of the
mouth,
pharynx and nasal cavities (
vocal tract) as the vocal fold vibration sound passes through producing varying resonant frequencies.
Central nervous control The analysis of brain lesions and the correlation between lesion locations and behavioral deficits were the most important sources of knowledge about the cerebral mechanisms underlying speech production for many years. The seminal lesion studies of
Paul Broca indicated that the production of speech relies on the functional integrity of the left
inferior frontal gyrus. More recently, the results of noninvasive neuroimaging techniques, such as
functional magnetic resonance imaging (fMRI), provide growing evidence that complex human skills are not primarily located in highly specialized brain areas (e.g.,
Broca's area) but are organized in networks connecting several different areas of both hemispheres instead.
Functional neuroimaging identified a complex neural network underlying speech production including cortical and subcortical areas, such as the
supplementary motor area,
cingulate motor areas,
primary motor cortex,
basal ganglia, and
cerebellum. ==Speech perception==