Caves provide a home for many unique biota. Cave ecologies are very diverse, and not sharply distinct from surface habitats. Generally however, the deeper the cave becomes, the more rarefied the ecology. Cave environments fall into three general categories: • Endogean :the parts of caves that are in communication with surface soils through cracks and rock seams, groundwater seepage, and root protrusion. • Parahypogean :the threshold regions near cave mouths that extend to the last penetration of sunlight. •
Hypogean :or "true" cave environments. These can be in regular contact with the surface via wind and underground rivers, or the migration of animals, or can be almost entirely isolated. Deep hypogean environments can host autonomous ecologies whose primary source of energy is not sunlight, but chemical energy liberated from limestone and other minerals by
chemoautotrophic bacteria. Cave organisms fall into three basic classes: There are also so-called
accidental trogloxenes which are surface organisms that enter caves for no survival reason. Some may even be
troglophobes (“cave haters”), which cannot survive in caves for any extended period. Examples include deer which fell through a sinkhole, frogs swept into a cave by a flash flood, etc. The two factors that limit cave ecologies are generally energy and nutrients. To some degree moisture is always available in actively forming Karst caves. Cut off from the sunlight and steady deposition of plant detritus, caves are poor habitats in comparison with wet areas on the surface. Most of the energy in cave environments comes from the surplus of the ecosystems outside. One major source of energy and nutrients in caves is dung from trogloxenes, most of which is deposited by bats. Other sources are mentioned above. Cave ecosystems are very fragile. Because of their rarity and position in the ecosystem they are threatened by a large number of human activities. Dam construction, limestone quarrying, water pollution and logging are just some of the disasters that can devastate or destroy underground biological communities. == Other areas of cave science ==