The single-player campaign spans the three eras of
Star Trek: Enterprise (ENT),
The Original Series (TOS) and
The Next Generation (TNG).
The Original Series era also includes elements from the
Original Series films.
The Next Generation era includes elements from
Deep Space Nine (DS9),
Voyager, (VOY) and the
Next Generation films. In each era, the player utilizes a fleet of Federation ships from that era to combat the Romulans (ENT), Klingons (TOS), and Borg (TNG). The eras covered in single-player, with the corresponding captains and main ships, are: •
Jonathan Archer (
Enterprise (NX-01) - ENT) •
James T. Kirk • (
USS Enterprise - TOS,
The Motion Picture,
The Wrath of Khan and
The Search for Spock) • (
USS Enterprise-A -
The Voyage Home,
The Final Frontier and
The Undiscovered Country) •
Jean-Luc Picard • (
USS Stargazer - TNG) • (
USS Enterprise-D - TNG and
Generations) • (
USS Enterprise-E -
First Contact,
Insurrection and
Nemesis) •
Benjamin Sisko (
USS Defiant - DS9) •
Kathryn Janeway (
USS Voyager - VOY)
Enterprise Era The story begins in 2159, approximately four years after the events of "
Demons" and "
Terra Prime" and two years before the decommissioning of
Enterprise (NX-01) in 2161 (see "
These Are the Voyages...") as Captain Jonathan Archer (voiced by
Scott Bakula) is sent to search for a missing
Vulcan science ship in a nearby star system.
Enterprise arrives but is immediately attacked by Romulans. After fighting off the Romulans,
Enterprise finds the Vulcan ship under attack from another group of Romulans. After saving the ship, the
Seleya, their captain, Commander T'Uerell (Bari Biern), asks
Enterprise to accompany her to her research station,
Gravenor, which, she fears, may also be under attack. Upon reaching the station, T'Uerell's fears are proven correct, as Romulans are attacking the station. T'Uerell docks to evacuate her people while
Enterprise holds off the incoming Romulans. However, T'Uerell breaks off radio contact and disembarks from the station seconds before the
Gravenor explodes, with a full crew complement still on board. As T'Uerell departs, she thanks Archer for defending her whilst she recovered her
protomatter from the station. She flees, and the badly damaged
Enterprise is forced to return to
Earth. Some time later,
Enterprise is ordered to escort three medical ships through a system where a
viral epidemic has broken out. Several times the Romulans attack, but each time, they are repulsed. The medical ships are able to deliver the
antidote, and the epidemic is cured. However, it is confirmed that the virus was artificially created and released into the planets'
ecosystem on purpose.
Enterprise is sent to discover where the virus came from. After identifying the design of a vessel seen navigating through the system just prior to the outbreak,
Enterprise proceeds to a nearby planet. Upon arrival, however, they find the virus has been released on the planet, and all life has been wiped out. They search through the system, finding several ships harvesting the virus from
gas giants and several harvesting stations orbiting nearby planets in preparation to release the virus into the atmosphere. They destroy the stations, but one harvesting ship is able to escape with a sample of the virus.
Enterprise then learns that the Romulans are mounting a large-scale attack near
Earth, and they arrive just in time to prevent the destruction of a starbase. They learn that the ship with the virus has had its contents distributed to three smaller ships, and these three ships are now being escorted towards Earth by the Romulan fleet. Using nearby mining facilities, which still use
nuclear fusion,
Enterprise is able to disable the engines of the three ships, which the starbase subsequently destroys. With this, the remaining Romulan ships disengage and Earth is saved.
Original Series Era The game then cuts to 2270, the era of the original
Constitution-class USS
Enterprise (NCC-1701) under the command of James T. Kirk (
William Shatner). It is the height of a war with the Klingon Empire, and Starfleet Intelligence has uncovered the development of an advanced Klingon weapons platform. Kirk is tasked with stealing an experimental Klingon Bird-of-Prey which has been fitted with a prototype cloaking device. After successfully doing so, Kirk pilots the vessel to the weapon platform. While under cloak, Kirk sees the
Seleya at a mining facility, and listens in on a conversation that reveals T'Uerell designed the weapons platform in exchange for mineral deposits. With this information in hand, Kirk recommends that Starfleet conduct an immediate strike on the platform. Kirk is assigned a task force and destroys the weapons platform, where he again encounters T'Uerell, who warps out of the system. Twenty years later, T'Uerell appears at a classified Federation research station. Now in command of the new
Constitution-class USS
Enterprise (NCC-1701-A), Admiral Kirk avoids going into retirement so as to deal with the matter. Kirk is able to regain control of the station after a skirmish with some Klingon vessels. Upon beaming an away team aboard, they report the presence of mysterious
cybernetic implants in what used to be the Klingons aboard the station. Kirk confronts T'Uerell, who again flees, leaving behind a large Borg sphere and three mysterious orbs. By destroying the three smaller spheres, Kirk is able to destroy the main sphere. However, upon the sphere's destruction, a massive explosion destroys subspace for a three light-year radius. As he contemplates his failure to capture T'Uerell, Kirk states, "Hopefully the next generation will do better."
Next Generation Era The game then cuts to 2333 as the USS
Stargazer monitors several Romulan vessels engaged in combat with T'Uerell in a nearby star system. The Romulans are destroyed when T'Uerell uses a super-weapon on a planet in the system, instantly liquefying it to molten rock and destabilizing its core. The shockwave from the weapon damages the
Stargazer, fatally wounding Captain Ruhalter (David Bryan Jackson) and advancing the young Jean-Luc Picard (
Patrick Stewart) to acting captain. The
Stargazer and several other Starfleet vessels successfully destroy most of the stellar debris emanating from the liquefied planet so as to prevent collisions with several inhabited planets nearby. The game then moves to 2368 as Commander Benjamin Sisko (
Avery Brooks), having recently been assigned command of the experimental USS
Defiant, takes his ship to the Itari system to test it out. En route, he encounters a mysterious phenomenon which prevents warp, along with several Romulan Warbirds. They intend to apprehend a dangerous fugitive allegedly responsible for the deaths of many Romulans. Sisko confronts the Romulans over their illegal presence in Federation space and is taken prisoner. Captain Jean-Luc Picard, now in command of the
Galaxy-class USS
Enterprise (NCC-1701-D), is sent to the system to locate and recover the
Defiant and her crew. After learning of the presence of warp inhibitors in the system, the
Enterprise destroys them and Picard's team assaults a Romulan starbase in the area, saving the
Defiant. The game then advances to 2380, following the end of the
Dominion War (see "
What You Leave Behind"), the return of the USS
Voyager from the
Delta Quadrant (see "
Endgame"), and the collapse of the Romulan Empire (see
Star Trek: Nemesis). Captain Picard and the
Sovereign-class USS
Enterprise (NCC-1701-E), along with Admiral Janeway (
Kate Mulgrew) aboard the USS
Voyager, are engaged in discussions with T'Uerell, who is escorted by several powerful Borg vessels. She reveals that she has assumed the role of the
Borg Queen, having reformed the Collective into a tool of total
logic in pursuit of her goals to eliminate all that is illogical in the galaxy. When Starfleet refuse to surrender, T'Uerell has the Borg begin to assimilate civilians on nearby planets. However, Picard and Janeway are able to co-ordinate a mass evacuation of the system. Meanwhile, several Romulan and Klingon fleets draw out T'Uerell's forces and lure it to the more defendable Deep Space 9 space station. The combined Federation, Klingon and Romulan fleets, assisted by Deep Space 9, then confront T'Uerell and her Borg forces. However, T'Uerell flees moments before her vessel can be destroyed. In another system, T'Uerell harnesses polaric ion energy from a
nebula to provide a massive boost to her shields. She then opens a transwarp conduit from the Delta Quadrant to bring in Borg reinforcements. With the battle raging, Picard devises a plan to disrupt the conduit by towing two sensor platforms modified to emit a magneton pulse to positions above and below T'Uerell's vessel. This pulse destabilizes the polaric ion energy string, disabling T'Uerell's ability to harness its energy for her shields and the conduit. With the transwarp conduit closed and her shields weakened, T'Uerell is finally defeated by the combined fleets of the Federation, Klingons and Romulans. ==Development==