Tropical rainforest

rain forest in Peru Tropical rainforests are hot and wet. Mean monthly temperatures exceed during all months of the year. Average annual rainfall is no less than and can exceed although it typically lies between and . This high level of precipitation often results in poor soils due to leaching of soluble nutrients in the ground. Tropical rainforests exhibit high levels of biodiversity. Around 40% to 75% of all biotic species are indigenous to the rainforests. Rainforests are home to half of all the living animal and plant species on the planet. Two-thirds of all flowering plants can be found in rainforests. A single hectare of rainforest may contain 42,000 different species of insect, up to 807 trees of 313 species and 1,500 species of higher plants. It is likely that there may be many millions of species of plants, insects and microorganisms still undiscovered in tropical rainforests. Tropical rainforests are among the most threatened ecosystems globally due to large-scale fragmentation as a result of human activity. Habitat fragmentation caused by geological processes such as volcanism and climate change occurred in the past, and have been identified as important drivers of speciation. However, fast human driven habitat destruction is suspected to be one of the major causes of species extinction. Tropical rain forests have been subjected to heavy logging and agricultural clearance throughout the 20th century, and the area covered by rainforests around the world is rapidly shrinking. History tree group has dominated the Borneo lowland rain forests for millions of years Tropical rainforests have existed on earth for hundreds of millions of years. Most tropical rainforests today are on fragments of the Mesozoic era supercontinent of Gondwana. The breakup of Gondwana left tropical rainforests located in five major regions of the world: tropical America, Africa, Southeast Asia, Madagascar, and New Guinea, with smaller outliers in Australia. However, the specifics of the origin of rainforests remain uncertain due to an incomplete fossil record. Other types of tropical forest Several biomes may appear similar to, or merge via ecotones with, tropical rainforest: seen from the Hana Highway ;Moist seasonal tropical forest: in Queensland is a seasonal tropical forest. Moist seasonal tropical forests receive high overall rainfall with a warm summer wet season and a cooler winter dry season. These forests usually fall under tropical monsoon or tropical savanna climates. Some trees in these forests drop some or all of their leaves during the winter dry season, thus they are sometimes called "tropical mixed forest". They are found in parts of South America, in Central America and around the Caribbean, in coastal West Africa, parts of the Indian subcontinent, and across much of Indochina. ;Montane rainforests: These are found in cooler-climate mountainous areas, becoming known as cloud forests at higher elevations. Depending on latitude, the lower limit of montane rainforests on large mountains is generally between 1500 and 2500m while the upper limit is usually from 2400 to 3300m. ;Flooded rainforests: Tropical freshwater swamp forests, or "flooded forests", are found in Amazon basin (the Várzea) and elsewhere. == Forest structure ==

Rainforests are divided into different strata, or layers, with vegetation organized into a vertical pattern from the top of the soil to the canopy. Forest floor The forest floor, the bottom-most layer, receives only 2% of the sunlight. Only plants adapted to low light can grow in this region. Away from riverbanks, swamps and clearings, where dense undergrowth is found, the forest floor is relatively clear of vegetation because of the low sunlight penetration. This more open quality permits the easy movement of larger animals such as: ungulates like the okapi (Okapia johnstoni), tapir (Tapirus sp.), Sumatran rhinoceros (Dicerorhinus sumatrensis), and apes like the western lowland gorilla (Gorilla gorilla), as well as many species of reptiles, amphibians, and insects. The forest floor also contains decaying plant and animal matter, which disappears quickly, because the warm, humid conditions promote rapid decay. Many forms of fungi growing here help decay the animal and plant waste. Understory layer The understory layer lies between the canopy and the forest floor. The understory is home to a number of birds, small mammals, insects, reptiles, and predators. Examples include leopard (Panthera pardus), poison dart frogs (Dendrobates sp.), ring-tailed coati (Nasua nasua), boa constrictor (Boa constrictor), and many species of Coleoptera. Some examples of emergents include: Hydrochorea elegans, Dipteryx panamensis, Hieronyma alchorneoides, Hymenolobium mesoamericanum, Lecythis ampla and Terminalia oblonga. These trees need to be able to withstand the hot temperatures and strong winds that occur above the canopy in some areas. Several unique faunal species inhabit this layer such as the crowned eagle (Stephanoaetus coronatus), the king colobus (Colobus polykomos), and the large flying fox (Pteropus vampyrus). However, stratification is not always clear. Rainforests are dynamic and many changes affect the structure of the forest. Emergent or canopy trees collapse, for example, causing gaps to form. Openings in the forest canopy are widely recognized as important for the establishment and growth of rainforest trees. It is estimated that perhaps 75% of the tree species at La Selva Biological Station, Costa Rica are dependent on canopy opening for seed germination or for growth beyond sapling size, for example. == Ecology ==

Climates , Mexico in French Guiana Tropical rainforests are located around and near the equator, therefore having what is called an equatorial climate characterized by three major climatic parameters: temperature, rainfall, and dry season intensity. Other parameters that affect tropical rainforests are carbon dioxide concentrations, solar radiation, and nitrogen availability. In general, climatic patterns consist of warm temperatures and high annual rainfall. However, the abundance of rainfall changes throughout the year creating distinct moist and dry seasons. Tropical forests are classified by the amount of rainfall received each year, which has allowed ecologists to define differences in these forests that look so similar in structure. According to Holdridge's classification of tropical ecosystems, true tropical rainforests have an annual rainfall greater than 2m and annual temperature greater than 24 degrees Celsius, with a potential evapotranspiration ratio (PET) value of <0.25. However, most lowland tropical forests can be classified as tropical moist or wet forests, which differ in regards to rainfall. Tropical forest ecology- dynamics, composition, and function- are sensitive to changes in climate especially changes in rainfall. Oxisols, infertile, deeply weathered and severely leached, have developed on the ancient Gondwanan shields. Rapid bacterial decay prevents the accumulation of humus. The concentration of iron and aluminium oxides by the laterization process gives the oxisols a bright red color and sometimes produces minable deposits (e.g., bauxite). On younger substrates, especially of volcanic origin, tropical soils may be quite fertile. Nutrient recycling This high rate of decomposition is the result of phosphorus levels in the soils, precipitation, high temperatures and the extensive microorganism communities. In addition to the bacteria and other microorganisms, there are an abundance of other decomposers such as fungi and termites that aid in the process as well. Nutrient recycling is important because below ground resource availability controls the above ground biomass and community structure of tropical rainforests. These soils are typically phosphorus limited, which inhibits net primary productivity or the uptake of carbon. The soil contains microbial organisms such as bacteria, which break down leaf litter and other organic matter into inorganic forms of carbon usable by plants through a process called decomposition. During the decomposition process the microbial community is respiring, taking up oxygen and releasing carbon dioxide. The decomposition rate can be evaluated by measuring the uptake of oxygen. High temperatures and precipitation increase decomposition rate, which allows plant litter to rapidly decay in tropical regions, releasing nutrients that are immediately taken up by plants through surface or ground waters. The seasonal patterns in respiration are controlled by leaf litter fall and precipitation, the driving force moving the decomposable carbon from the litter to the soil. Respiration rates are highest early in the wet season because the recent dry season results in a large percentage of leaf litter and thus a higher percentage of organic matter being leached into the soil. Buttress roots '' in eastern Ecuador A common feature of many tropical rainforests is the distinct buttress roots of trees. Instead of penetrating to deeper soil layers, buttress roots create a widespread root network at the surface for more efficient uptake of nutrients in a very nutrient poor and competitive environment. Most of the nutrients within the soil of a tropical rainforest occur near the surface because of the rapid turnover time and decomposition of organisms and leaves. Because of this, the buttress roots occur at the surface so the trees can maximize uptake and actively compete with the rapid uptake of other trees. These roots also aid in water uptake and storage, increase surface area for gas exchange, and collect leaf litter for added nutrition. Additionally, these roots reduce soil erosion and maximize nutrient acquisition during heavy rains by diverting nutrient rich water flowing down the trunk into several smaller flows while also acting as a barrier to ground flow. Also, the large surface areas these roots create provide support and stability to rainforests trees, which commonly grow to significant heights. This added stability allows these trees to withstand the impacts of severe storms, thus reducing the occurrence of fallen trees. Forest succession Succession is an ecological process that changes the biotic community structure over time towards a more stable, diverse community structure after an initial disturbance to the community. The initial disturbance is often a natural phenomenon or human caused event. Natural disturbances include hurricanes, volcanic eruptions, river movements or an event as small as a fallen tree that creates gaps in the forest. In tropical rainforests, these same natural disturbances have been well documented in the fossil record, and are credited with encouraging speciation and endemism. Human land use practices have led to large-scale deforestation. In many tropical countries such as Costa Rica these deforested lands have been abandoned and forests have been allowed to regenerate through ecological succession. These regenerating young successional forests are called secondary forests or second-growth forests. == Biodiversity and speciation ==

at Bukit Lawang, Sumatra Tropical rainforests exhibit a vast diversity in plant and animal species. The root for this remarkable speciation has been a query of scientists and ecologists for years. A number of theories have been developed for why and how the tropics can be so diverse. Interspecific competition Interspecific competition results from a high density of species with similar niches in the tropics and limited resources available. Species which "lose" the competition may either become extinct or find a new niche. Direct competition will often lead to one species dominating another by some advantage, ultimately driving it to extinction. Niche partitioning is the other option for a species. This is the separation and rationing of necessary resources by utilizing different habitats, food sources, cover or general behavioral differences. A species with similar food items but different feeding times is an example of niche partitioning. Pleistocene refugia The theory of Pleistocene refugia was developed by Jürgen Haffer in 1969 with his article Speciation of Amazonian Forest Birds. Haffer proposed the explanation for speciation was the product of rainforest patches being separated by stretches of non-forest vegetation during the last glacial period. He called these patches of rainforest areas refuges and within these patches allopatric speciation occurred. With the end of the glacial period and increase in atmospheric humidity, rainforest began to expand and the refuges reconnected. This theory has been the subject of debate. Scientists are still skeptical of whether or not this theory is legitimate. Genetic evidence suggests speciation had occurred in certain taxa 1–2 million years ago, preceding the Pleistocene. == Human dimensions ==

Habitation in New Guinea Tropical rainforests have harboured human life for many millennia, with many Indigenous people in South and Central America, who belong to the Indigenous peoples of the Americas, the Congo Pygmies in Central Africa, and several tribes in Southeast Asia, like the Dayak people and the Penan people in Borneo. On 18 January 2007, FUNAI reported also that it had confirmed the presence of 67 different uncontacted tribes in Brazil, up from 40 in 2005. With this addition, Brazil has now overtaken the island of New Guinea as the country having the largest number of uncontacted tribes. The province of Irian Jaya or West Papua in the island of New Guinea is home to an estimated 44 uncontacted tribal groups. hunter-gatherers in the Congo Basin hunters in Ecuador The pygmy peoples are hunter-gatherer groups living in equatorial rainforests characterized by their short height (below one and a half meters, or 59inches, on average). Amongst this group are the Efe, Aka, Twa, Baka, and Mbuti people of Central Africa. However, the term pygmy is considered pejorative so many tribes prefer not to be labeled as such. Some notable indigenous peoples of the Americas, or Amerindians, include the Huaorani, Ya̧nomamö, and Kayapo people of the Amazon. The traditional agricultural system practiced by tribes in the Amazon is based on swidden cultivation (also known as slash-and-burn or shifting cultivation) and is considered a relatively benign disturbance. Pygmies who live in Southeast Asia are, amongst others, referred to as "Negrito". Resources Cultivated foods and spices Yam, coffee, chocolate, banana, mango, papaya, macadamia, avocado, and sugarcane all originally came from tropical rainforest and are still mostly grown on plantations in regions that were formerly primary forest. In the mid-1980s and 1990s, 40 million tons of bananas were consumed worldwide each year, along with 13 million tons of mango. Central American coffee exports were worth US$3 billion in 1970. Much of the genetic variation used in evading the damage caused by new pests is still derived from resistant wild stock. Tropical forests have supplied 250 cultivated kinds of fruit, compared to only 20 for temperate forests. Forests in New Guinea alone contain 251 tree species with edible fruits, of which only 43 had been established as cultivated crops by 1985. Ecosystem services In addition to extractive human uses, rain forests also have non-extractive uses that are frequently summarized as ecosystem services. Rain forests play an important role in maintaining biological diversity, sequestering and storing carbon, global climate regulation, disease control, and pollination. Deforestation in the Amazon rainforest region was one of the main reason that cause the severe Drought of 2014–2015 in Brazil For the last three decades, the amount of carbon absorbed by the world's intact tropical forests has fallen, according to a study published in 2020 in the journal Nature. In 2019 they took up a third less carbon than they did in the 1990s, due to higher temperatures, droughts and deforestation. The typical tropical forest may become a carbon source by the 2060s. Tourism for seeing the diverse tropical forest in Costa Rica Despite the negative effects of tourism in the tropical rainforests, there are also several important positive effects. • In recent years ecotourism in the tropics has increased. While rainforests are becoming increasingly rare, people are travelling to nations that still have this diverse habitat. Locals are benefiting from the additional income brought in by visitors, as well areas deemed interesting for visitors are often conserved. Ecotourism can be an incentive for conservation, especially when it triggers positive economic change. == Conservation ==

forest in the tropics has varied from year to year, but remains consistently higher than preceding decades. Threats Deforestation in Brazil's Maranhão state, 2016 Mining and drilling in southwestern Papua New Guinea Deposits of precious metals (gold, silver, coltan) and fossil fuels (oil and natural gas) occur underneath rainforests globally. These resources are important to developing nations and their extraction is often given priority to encourage economic growth. Mining and drilling can require large amounts of land development, directly causing deforestation. In Ghana, a West African nation, deforestation from decades of mining activity left about 12% of the country's original rainforest intact. Conversion to agricultural land With the invention of agriculture, humans were able to clear sections of rainforest to produce crops, converting it to open farmland. Such people, however, obtain their food primarily from farm plots cleared from the forest and hunt and forage within the forest to supplement this. The issue arising is between the independent farmer providing for his family and the needs and wants of the globe as a whole. This issue has seen little improvement because no plan has been established for all parties to be aided. As major carbon reducers and carbon and soil methane storages, their destruction contributes to increasing global energy trapping, atmospheric gases. Climate change has been significantly contributed to by the destruction of the rainforests. A simulation was performed in which all rainforest in Africa were removed. The simulation showed an increase in atmospheric temperature by 2.5 to 5 degrees Celsius. The seasonality of rainforests affects the reproductive patterns of amphibians, and this in turn can directly affect the species of reptiles that feed on these groups, particularly species with specialized feeding, since these are less likely to use alternative resources. Protection Efforts to protect and conserve tropical rainforest habitats are diverse and widespread. Tropical rainforest conservation ranges from strict habitat preservation to finding sustainable management techniques for people living in tropical rainforests. International policy has also introduced a market incentive program called Reducing Emissions from Deforestation and Forest Degradation (REDD) for companies and governments to outset their carbon emissions through financial investments into rainforest conservation. == See also ==