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Phylum

In biology, a phylum is a level of classification, or taxonomic rank, that is below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 32 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about eight phyla. Current research in phylogenetics is uncovering the relationships among phyla within larger clades like Ecdysozoa and Embryophyta.

General description
The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, 'race, stock'), related to (, 'tribe, clan'). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained unity"): "perhaps such a real and completely self-contained unity is the aggregate of all species which have gradually evolved from one and the same common original form, as, for example, all vertebrates. We name this aggregate [a] [i.e., stock / tribe] ()." In plant taxonomy, August W. Eichler (1883) classified plants into five groups named divisions, a term that remains in use today for groups of plants, algae and fungi. The definitions of zoological phyla have changed from their origins in the six Linnaean classes and the four of Georges Cuvier. At its most basic, a phylum can be defined in two ways: as a group of organisms with a certain degree of morphological or developmental similarity (the phenetic definition), or a group of organisms with a certain degree of evolutionary relatedness (the phylogenetic definition). Attempting to define a level of the Linnean hierarchy without referring to (evolutionary) relatedness is unsatisfactory, but a phenetic definition is useful when addressing questions of a morphological nature—such as how successful different body plans were. Definition based on genetic relation The most important objective measure in the above definitions is the "certain degree" that defines how different organisms need to be members of different phyla. The minimal requirement is that all organisms in a phylum should be clearly more closely related to one another than to any other group. On the other hand, the highly parasitic phylum Mesozoa was divided into two phyla (Orthonectida and Rhombozoa) when it was discovered the Orthonectida are probably deuterostomes and the Rhombozoa protostomes. This changeability of phyla has led some biologists to call for the concept of a phylum to be abandoned in favour of placing taxa in clades without any formal ranking of group size. A classification using this definition may be strongly affected by the chance survival of rare groups, which can make a phylum much more diverse than it would be otherwise. == Known phyla ==
Known phyla
Animals Total numbers are estimates; figures from different authors vary wildly, not least because some are based on described species, and some on extrapolations to numbers of undescribed species. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of the total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million. Plants The kingdom Plantae is defined in various ways by different biologists (see Current definitions of Plantae). All definitions include the living embryophytes (land plants), to which may be added the two green algae divisions, Chlorophyta and Charophyta, to form the clade Viridiplantae. The table below follows the influential (though contentious) Cavalier-Smith system in equating "Plantae" with Archaeplastida, a group containing Viridiplantae and the algal Rhodophyta and Glaucophyta divisions. The definition and classification of plants at the division level also varies from source to source, and has changed progressively in recent years. Thus some sources place horsetails in division Arthrophyta and ferns in division Monilophyta, while others place them both in Monilophyta, as shown below. The division Pinophyta may be used for all gymnosperms (i.e. including cycads, ginkgos and gnetophytes), or for conifers alone as below. Since the first publication of the APG system in 1998, which proposed a classification of angiosperms up to the level of orders, many sources have preferred to treat ranks higher than orders as informal clades. Where formal ranks have been provided, the traditional divisions listed below have been reduced to a very much lower level, e.g. subclasses. Fungi Phylum Microsporidia is generally included in kingdom Fungi, though its exact relations remain uncertain, and it is considered a protozoan by the International Society of Protistologists which is considered undesirable by many biologists. Accordingly, there is a proposal to abolish the Zygomycota phylum. Its members would be divided between phylum Glomeromycota and four new subphyla incertae sedis (of uncertain placement): Entomophthoromycotina, Kickxellomycotina, Mucoromycotina, and Zoopagomycotina. which is less acceptable to present-day biologists than in the past. Proposals have been made to divide it among several new kingdoms, such as Protozoa and Chromista in the Cavalier-Smith system. Protist taxonomy has long been unstable, with different approaches and definitions resulting in many competing classification schemes. Many of the phyla listed below are used by the Catalogue of Life, and correspond to the Protozoa-Chromista scheme, with updates from the latest (2022) publication by Cavalier-Smith. Other phyla are used commonly by other authors, and are adapted from the system used by the International Society of Protistologists (ISP). Some of the descriptions are based on the 2019 revision of eukaryotes by the ISP. The number of protist phyla varies greatly from one classification to the next. The Catalogue of Life includes Rhodophyta and Glaucophyta in kingdom Plantae, In addition, less popular classification schemes unite Ochrophyta and Pseudofungi under one phylum, Gyrista, and all alveolates except ciliates in one phylum Myzozoa, later lowered in rank and included in a paraphyletic phylum Miozoa. • AbditibacteriotaAcidobacteriota, phenotypically diverse and mostly uncultured • Actinomycetota, High-G+C Gram positive species • Aquificota, deep-branching • ArmatimonadotaAtribacterotaBacillota, Low-G+C Gram positive species, such as the spore-formers Bacilli (aerobic) and Clostridia (anaerobic) • BacteroidotaBalneolotaBdellovibrionotaCaldisericota, formerly candidate division OP5, Caldisericum exile is the sole representative • CalditrichotaCampylobacterotaChlamydiotaChlorobiota, green sulphur bacteria • Chloroflexota, green non-sulphur bacteria • Chrysiogenota, only 3 genera (Chrysiogenes arsenatis, Desulfurispira natronophila, Desulfurispirillum alkaliphilum) • CoprothermobacterotaDeferribacterotaDeinococcota, Deinococcus radiodurans and Thermus aquaticus are "commonly known" species of this phyla • DictyoglomotaElusimicrobiota, formerly candidate division Thermite Group 1 • FibrobacterotaFusobacteriotaGemmatimonadotaIgnavibacteriotaKiritimatiellotaLentisphaerota, formerly clade VadinBE97 • Mycoplasmatota, notable genus: MycoplasmaMyxococcotaNitrospinotaNitrospirotaPlanctomycetotaPseudomonadota, the most well-known phylum, containing species such as Escherichia coli or Pseudomonas aeruginosaRhodothermotaSpirochaetota, species include Borrelia burgdorferi, which causes Lyme disease • SynergistotaThermodesulfobacteriotaThermomicrobiotaThermotogota, deep-branching • Verrucomicrobiota Archaea Currently there are 2 phyla that have been validly published according to the Bacteriological CodeNitrososphaerotaThermoproteota, second most common archaeal phylum Other phyla that have been proposed, but not validly named, include: • "Euryarchaeota", most common archaeal phylum • "Korarchaeota" • "Nanoarchaeota", ultra-small symbiotes, single known species == See also ==
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