The Leptoconopinae is a
subfamily of biting midges. The larvae are recognized by their unique
sclerites of the head, and by their mouthparts. The Forcipomyiinae are a subfamily of biting midges. In this subfamily, both anterior and posterior
prolegs are present on the larvae. Larvae are both terrestrial and aquatic, and feed primarily on algae and fungi. Some species are important
pollinators of tropical crops such as the
cocoa bean.
Systematics Basal lineages • †
Lebanoculicoides Szadziewski 1996
Lebanese amber,
Barremian,
Spanish amber,
Albian • Subfamily Leptoconopinae Noe 1907 • †
Archiaustroconops Szadziewski 1996
Durlston Formation, United Kingdom, Berriasian, Lebanese amber, Barremian,
Jordanian amber, Spanish amber, Albian,
Burmese amber, Myanmar,
Cenomanian •
Austroconops Wirth and Lee 1958 Barremian–Present • †
Fossileptoconops Szadziewski 1996 Lebanese amber, Barremian • †
Jordanoconops Szadziewski 2000 Jordanian amber, Albian •
Leptoconops Skuse 1889 Barremian–Present • †
Minyohelea Borkent 1995
Austrian amber,
Hauterivian Lebanese amber, Barremian,
Canadian amber,
Campanian • †
Archiculicoides Szadziewski 1996 Lebanese amber, Barremian • †
Gerontodacus Borkent 2019 Lebanese amber, Barremian, Spanish amber, Albian • †
Protoculicoides Boesel 1937 Burmese amber, Myanmar, Cenomanian, Canadian amber, Campanian • †
Atriculicoides Remm 1976 Spanish amber, Albian
Durtal amber, France, Cenomanian, Burmese amber, Myanmar, Cenomanian,
Taimyr amber, Russia, Cenomanian,
Santonian •
Dasyhelea Kieffer 1913 • Subfamily Forcipomyiinae •
Atrichopogon Kieffer 1906 •
Forcipomyia Meigen, 1818 • †
Adelohelea Borkent 1995
Hungarian amber, Santonian, Canadian amber, Campanian • †
Heleageron Borkent 1995
New Jersey amber,
Turonian Canadian amber, Campanian • †
Alautunmyia Borkent 1996
New Jersey amber,
Turonian ==Gallery==