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Astrothelium

Astrothelium is a large genus of corticolous (bark-dwelling) lichens in the family Trypetheliaceae. The genus is characterized by a corticate thallus and diverse ascomata structures, which can be simple, aggregated, or forming pseudostromata. Astrothelium is also notable for the carbonized walls of its ascomata, the so-called textura intricata arrangement of cells in these walls, and various forms of distoseptate, transparent spores.

Taxonomy
The genus was circumscribed in 1824 by German botanist Franz Gerhard Eschweiler, with Astrothelium conicum assigned as the type species. ==Description==
Description
The thallus, or the vegetative body of Astrothelium, is , meaning it is covered by a protective . The ascomata, which are the fruiting bodies containing the reproductive spores, can appear in various forms such as simple, aggregated, or forming . These structures often differ in their composition and colour, and can be used as to distinguish between species. The ascomata can range from being immersed within the thallus to prominently displayed on the surface. The ostioles, or small openings that allow the release of spores, can be located at the apex or eccentrically on the ascomata and may appear as either simple or fused structures. Astrothelium is also notable for its ascomatal wall, which is made up of hyphal cells and is typically carbonized. The wall features a , a term used to describe a tightly interwoven arrangement of cells. Within the , the tissue that houses the asci (spore-producing structures), the cells are either clear or filled with oil droplets. The ascospores, which are the sexual spores responsible for reproduction, are and hyaline in appearance. They can also exhibit a variety of forms, including transversely septate or , reflecting their segmented or multi-cellular nature. ==Species==
Species
, Species Fungorum accepts 267 species in Astrothelium. • Astrothelium aenascensAstrothelium aeneoides – Brazil • Astrothelium aeneumAstrothelium alboverrucoidesAstrothelium alboverrucumAstrothelium amazonumAstrothelium ambiguumAstrothelium amylosporum – Bolivia • Astrothelium andamanicumAstrothelium annulareAstrothelium astrolucidum – Brazil • Astrothelium aurantiacocinereumAstrothelium aurantiacumAstrothelium aurantioseptemseptatum – Brazil • Astrothelium auratumAstrothelium aureoirregulareAstrothelium aureomaculatumAstrothelium basilicumAstrothelium bicolorAstrothelium bireagens – Colombia • Astrothelium bivelum – Brazil • Astrothelium buckiiAstrothelium bulbosum – Brazil • Astrothelium bullatothallinum – Venezuela • Astrothelium bullatum – Bolivia • Astrothelium calosporumAstrothelium campylocartilagineumAstrothelium carassenseAstrothelium carrascoense – Bolivia • Astrothelium cartilagineumAstrothelium cayennense – French Guiana • Astrothelium cecidiogenumAstrothelium ceratinumAstrothelium chapadenseAstrothelium chrysoglyphumAstrothelium chrysostomumAstrothelium chulumanense – Bolivia • Astrothelium cinereorosellumAstrothelium cinereumAstrothelium citrisporum – Brazil • Astrothelium clypeatumAstrothelium coccineum – Mexico • Astrothelium colombienseAstrothelium coloratum – Brazil • Astrothelium condoricumAstrothelium confluensAstrothelium conicumAstrothelium conjugatumAstrothelium consimileAstrothelium corallinumAstrothelium corticatumAstrothelium crassumAstrothelium cryptolucensAstrothelium curvatum – Brazil • Astrothelium curvisporum – Brazil • Astrothelium decemseptatum – Brazil • Astrothelium deformeAstrothelium defossumAstrothelium degeneransAstrothelium diaphanocorticatum – Papua New Guinea • Astrothelium dicoloratumAstrothelium dimidioinspersum – Brazil • Astrothelium disjunctum – Brazil • Astrothelium dissimilumAstrothelium duplicatum – Brazil • Astrothelium ecuadorenseAstrothelium effusumAstrothelium elixii – Bolivia • Astrothelium endochryseumAstrothelium eumultiseptatum – Brazil • Astrothelium eustominspersum – Brazil • Astrothelium eustomurale – Brazil • Astrothelium exostemmatisAstrothelium fallaxAstrothelium feeiAstrothelium fernandaeAstrothelium ferrugineumAstrothelium fijienseAstrothelium flavocoronatumAstrothelium flavoduplex – Brazil • Astrothelium flavogigasporum – Brazil • Astrothelium flavomaculatumAstrothelium flavomegaspermumAstrothelium flavomeristosporumAstrothelium flavomurisporum – Brazil • Astrothelium flavostiolatumAstrothelium flavostromatum – Brazil • Astrothelium flavum – Brazil • Astrothelium fuscosporumAstrothelium galbineumAstrothelium galligenumAstrothelium gigantosporumAstrothelium globosum – Brazil • Astrothelium graphicum – Brazil • Astrothelium grossoidesAstrothelium guianenseAstrothelium gyalostiolatum – Brazil • Astrothelium himalayenseAstrothelium indicumAstrothelium infossumAstrothelium infravulcanum – Brazil • Astrothelium infuscatulumAstrothelium inspersaeneumAstrothelium inspersonitidulum – Brazil • Astrothelium inspersoconicumAstrothelium inspersogalbineumAstrothelium inspersonovemseptatum – Brazil • Astrothelium inspersotuberculosum – Bolivia • Astrothelium insulare – Brazil • Astrothelium intermediumAstrothelium introflavidum – Brazil • Astrothelium irregulareAstrothelium isidiatum – Bolivia • Astrothelium isohypocrellinumAstrothelium jiangxienseAstrothelium keralenseAstrothelium komposchiiAstrothelium kunzeiAstrothelium laevithallinumAstrothelium laurerosphaerioidesAstrothelium lauroides – Brazil • Astrothelium leioplacumAstrothelium leucosessileAstrothelium lineatumAstrothelium longisporum – Brazil • Astrothelium lucidomedullatumAstrothelium lucidostromumAstrothelium lucidothallinumAstrothelium lugescensAstrothelium luminothallinumAstrothelium luridumAstrothelium macrocarpumAstrothelium macroeustomum – French Guiana • Astrothelium macrosporumAstrothelium macrostiolatumAstrothelium macrostomoidesAstrothelium macrostomum – Brazil • Astrothelium marcidumAstrothelium marjoleinae – Brazil • Astrothelium meandratum – Brazil • Astrothelium mediocrassumAstrothelium medioincrassatum – Brazil • Astrothelium megacrypticumAstrothelium megaeneum – Bolivia • Astrothelium megaleiumAstrothelium megalophthalmumAstrothelium megalostomumAstrothelium megaspermumAstrothelium megatropicumAstrothelium megeustomum – Brazil • Astrothelium megeustomurale – Brazil • Astrothelium meghalayenseAstrothelium megochroleucumAstrothelium meiophorumAstrothelium meristosporoidesAstrothelium meristosporumAstrothelium mesoduplex – Brazil • Astrothelium miniannulareAstrothelium minicecidiogenum – Costa Rica • Astrothelium mordonialenseAstrothelium multireflexum – Brazil • Astrothelium muriconicumAstrothelium myopicum – Brazil • Astrothelium neglectumAstrothelium neodiplocarpum – Bolivia • Astrothelium neogalbineumAstrothelium neoinspersumAstrothelium neovariolosumAstrothelium nicaraguenseAstrothelium nigratumAstrothelium nigrocacuminum – Bolivia • Astrothelium nigrorufumAstrothelium nigrum – Brazil • Astrothelium nitidiusculumAstrothelium nitidulumAstrothelium norisianumAstrothelium novemseptatum – Brazil • Astrothelium obtectumAstrothelium ochraceumAstrothelium ochroleucoides – Brazil • Astrothelium octoseptatum – Brazil • Astrothelium octosporoidesAstrothelium octosporumAstrothelium oligocarpumAstrothelium olivaceofuscumAstrothelium palaeoexostemmatis – Thailand • Astrothelium pallidoflavum – Bolivia • Astrothelium papillosumAstrothelium papulosumAstrothelium parabathelium – Brazil • Astrothelium parathelioides – Brazil • Astrothelium perancepsAstrothelium perspersumAstrothelium phaeotheliumAstrothelium philippinenseAstrothelium phlyctaenaAstrothelium pictum – Brazil • Astrothelium porosumAstrothelium praetervisumAstrothelium pseudannulareAstrothelium pseudocrassumAstrothelium pseudodermatodesAstrothelium pseudodissimulumAstrothelium pseudoferrugineumAstrothelium pseudomegalophthalmumAstrothelium pseudoplatystomumAstrothelium pseudovariatumAstrothelium puiggariiAstrothelium pulcherrimumAstrothelium pupulaAstrothelium purpurascensAstrothelium pustulatumAstrothelium pyrenuliforme – Bolivia • Astrothelium quasimamillanum – Brazil • Astrothelium quatuorseptatum – Brazil • Astrothelium quintannulare – Brazil • Astrothelium quintosulphureum – Brazil • Astrothelium rhinothallinumAstrothelium rimosumAstrothelium robustosporum – Brazil • Astrothelium rogitamaeAstrothelium rubrocrystallinum – Brazil • Astrothelium rubrostiolatum – Brazil • Astrothelium rufescensAstrothelium sanguinariumAstrothelium sanguineoxanthumAstrothelium santessoniiAstrothelium saxicolaAstrothelium scoriaAstrothelium scoriotheliumAstrothelium scorizumAstrothelium septemseptatumAstrothelium septoconicumAstrothelium sexloculatumAstrothelium siamenseAstrothelium sierraleonenseAstrothelium sikkimenseAstrothelium simplex – Brazil • Astrothelium sinenseAstrothelium sinuosum – Brazil • Astrothelium sipmaniiAstrothelium solitarium – Brazil • Astrothelium sorditheciumAstrothelium spectabileAstrothelium sphaerioidesAstrothelium squamosum – Brazil • Astrothelium stellare – Brazil • Astrothelium stramineumAstrothelium straminicolorAstrothelium stromatocinnamomeum – Brazil • Astrothelium stromatofluorescens – Brazil • Astrothelium stromatolucidum – Colombia • Astrothelium studerae – Brazil • Astrothelium subcatervariumAstrothelium subdiscretumAstrothelium subdisjunctumAstrothelium subdissociansAstrothelium subendochryseumAstrothelium subeustominspersumAstrothelium subinterjectumAstrothelium subnitidiusculumAstrothelium subrufescensAstrothelium subscoria – Bolivia • Astrothelium subsiamenseAstrothelium sulphureumAstrothelium superbumAstrothelium supraclandestinum – Brazil • Astrothelium suprainspersum – Brazil • Astrothelium tanianum – Malaysia • Astrothelium tenueAstrothelium testudineum – Brazil • Astrothelium tetrasporum – Brazil • Astrothelium thelotremoidesAstrothelium trematumAstrothelium trypethelioidesAstrothelium trypethelizansAstrothelium tuberculosumAstrothelium ubianenseAstrothelium ultralucensAstrothelium unisporumAstrothelium valsoides – Brazil • Astrothelium variabile – Bolivia • Astrothelium variatumAstrothelium vezdaeAstrothelium vulcanumAstrothelium xanthocavatum – Brazil • Astrothelium xanthopseudocyphellatumAstrothelium xanthosordithecium – Brazil • Astrothelium xanthosuperbum – Brazil • Astrothelium zebrinum ==References==
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