Autotroph

The term autotroph was coined by the German botanist Albert Bernhard Frank in 1892. It stems from the ancient Greek word (), meaning "nourishment" or "food". The first autotrophic organisms likely evolved early in the Archean but proliferated across Earth's Great Oxidation Event with an increase to the rate of oxygenic photosynthesis by cyanobacteria. Photoautotrophs evolved from heterotrophic bacteria by developing photosynthesis. The earliest photosynthetic bacteria used hydrogen sulfide. Due to the scarcity of hydrogen sulfide, some photosynthetic bacteria evolved to use water in photosynthesis, leading to cyanobacteria. ==Variants==

Some organisms rely on organic compounds as a source of carbon, but are able to use light or inorganic compounds as a source of energy. Such organisms are mixotrophs. An organism that obtains carbon from organic compounds but obtains energy from light is called a photoheterotroph, while an organism that obtains carbon from organic compounds and energy from the oxidation of inorganic compounds is termed a chemolithoheterotroph. Evidence suggests that some fungi may also obtain energy from ionizing radiation: Such radiotrophic fungi were found growing inside a reactor of the Chernobyl nuclear power plant. ==Examples==

There are many different types of autotrophs in Earth's ecosystems. Lichens located in tundra climates are an exceptional example of a primary producer that, by mutualistic symbiosis, combines photosynthesis by algae (or additionally nitrogen fixation by cyanobacteria) with the protection of a decomposer fungus. As there are many examples of primary producers, two dominant types are coral and one of the many types of brown algae, kelp. ==Photosynthesis==

Gross primary production occurs by photosynthesis. This is the main way that primary producers get energy and make it available to other forms of life. Plants, many corals (by means of intracellular algae), some bacteria (cyanobacteria), and algae do this. During photosynthesis, primary producers receive energy from the sun and use it to produce sugar and oxygen. ==Ecology==

, a photoautotroph Without primary producers, organisms that are capable of producing energy on their own, the biological systems of Earth would be unable to sustain themselves. The process of photosynthesis splits a water molecule (H2O), releasing oxygen (O2) into the atmosphere, and reducing carbon dioxide (CO2) to release hydrogen atoms that fuel the metabolic process of primary production. Plants convert and store the energy of the photons into the chemical bonds of simple sugars during photosynthesis. These plant sugars are polymerized for storage as long-chain carbohydrates, such as starch and cellulose; glucose is also used to make fats and proteins. When autotrophs are eaten by heterotrophs, i.e., consumers such as animals, the carbohydrates, fats, and proteins contained in them become energy sources for the heterotrophs. Proteins can be made using nitrates, sulfates, and phosphates in the soil. Primary production in tropical streams and rivers Aquatic algae are a significant contributor to food webs in tropical rivers and streams. This is displayed by net primary production, a fundamental ecological process that reflects the amount of carbon that is synthesized within an ecosystem. This carbon ultimately becomes available to consumers. Net primary production displays that the rates of in-stream primary production in tropical regions are at least an order of magnitude greater than in similar temperate systems. ==Origin of autotrophs==

Researchers believe that the first cellular lifeforms were not heterotrophs as they would rely upon autotrophs since organic substrates delivered from space were either too heterogeneous to support microbial growth or too reduced to be fermented. Instead, they consider that the first cells were autotrophs. These autotrophs might have been thermophilic and anaerobic chemolithoautotrophs that lived at deep sea alkaline hydrothermal vents. This view is supported by phylogenetic evidencethe physiology and habitat of the last universal common ancestor (LUCA) is inferred to have also been a thermophilic anaerobe with a Wood-Ljungdahl pathway, its biochemistry was replete with FeS clusters and radical reaction mechanisms. It was dependent upon Fe, H2, and CO2. The high concentration of K+ present within the cytosol of most life forms suggests that early cellular life had Na+/H+ antiporters or possibly symporters. Autotrophs possibly evolved into heterotrophs when they were at low H2 partial pressures where the first form of heterotrophy were likely amino acid and clostridial type purine fermentations. It has been suggested that photosynthesis emerged in the presence of faint near-infrared light emitted by hydrothermal vents. The first photochemically active pigments are then thought to be Zn-tetrapyrroles. ==See also==