Species within this genus have been found in the
Eocene of the
United States, in the
Paleocene of
Colombia and the
United States, and in the
Cretaceous of
Angola,
Argentina,
Brazil,
Cameroon,
Egypt,
France,
Iran,
Jordan,
Libya,
Morocco,
Mozambique,
Niger,
Nigeria,
Oman,
Pakistan,
Saudi Arabia,
Senegal and
Tunisia. ==References==