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Bignoniaceae

Bignoniaceae is a family of flowering plants in the order Lamiales commonly known as the bignonias or trumpet vines. It is not known to which of the other families in the order it is most closely related.

Description
'' Members of this family are mostly trees or lianas, sometimes shrubs, and rarely subshrubs or herbs. Lianas of the tribe Bignonieae have a unique vascular structure, in which phloem arms extend downward into the xylem because certain segments of the cambium cease the production of xylem at an early stage of development. The number of these arms is four or a multiple thereof, up to 32. The leaves are petiolate. Leaf arrangement usually is opposite, or rarely alternate or verticillate (in whorls). Leaves are usually compound, bifoliate, trifoliate, pinnate, or palmate, or rarely simple. Stipules are absent, but persistent; enlarged axillary bud scales (pseudostipules) are often present. Domatia occur in some genera. '' in Hyderabad, India Flowers are solitary or in inflorescences in a raceme or a helicoid or dichasial cyme. Inflorescences bear persistent or deciduous bracts or bractlets. The flowers are hypogynous, zygomorphic, bisexual, and usually conspicuous. The calyx and corolla are distinct. The calyx is synsepalous, with five sepals. The corolla is sympetalous, with five petals, often bilabiate. Corolla lobes are imbricate in bud, or rarely valvate, and usually much shorter than the corolla tube. Stamens are inserted on the corolla tube, alternating with corolla lobes. The four stamens are didynamous, members of each pair often connivent, the adaxial stamen is usually staminodial or absent; rarely with five fertile stamens or with two fertile and three staminodial stamens. The stigma is bilobed, and usually sensitive; a style is present. The ovary is superior, usually surrounded by a nectary disk, composed of two carpels, bilocular and with a septum, except unilocular in Tourrettia and quadrilocular in Eccremocarpus. Placentation is axile, except parietal in Tourrettia. Ovules are numerous. 4 stamens and the style-stigma, all in dorsal position. The fruit is usually a bivalved capsule, often with a replum. Dehiscence is septicidal or loculicidal. The three exceptions are the genera Kigelia, Crescentia and their close relatives, and Colea and its close relatives. In these, the fruit is indehiscent, not a capsule, and the seeds are not winged. The fruit is a berry in these genera. Seeds are usually flat and winged. Aril is absent. Endosperm usually absent, and sometimes sparse. Iridoids are usually present. Other compounds detected in Bignoniaceae include verbascosides, cornoside, quercetin, ursolic acid, saponins, and catalpic acid. The chromosome number does not vary much in Bignoniaceae. The haploid (base chromosome number) is 20 for nearly every species sampled, but some species have very small chromosomes, making an accurate count difficult. B chromosomes are common in Bignoniaceae. Pollination is either entomophilous (via insects), ornithophilous (via birds), or chiropterophilous (via bats). == Taxonomy ==
Taxonomy
The family Bignoniaceae was first validly published in the botanical literature (as Bignonieae) by Antoine Laurent de Jussieu in 1789 in his classic work, Genera Plantarum. The type genus for this family is Bignonia, which was validated by Linnaeus in Species Plantarum in 1753. The name originated with Joseph Pitton de Tournefort, who named it for his benefactor, Jean-Paul Bignon, in 1694, in his influential Eléments de botanique ou méthode pour connaître les plantes. Important groundwork for future study of the family was laid down from 1789 to 1837, mostly by Jussieu, Kunth, Bojer and G.Don (George Don (1798–1856) not George Don the elder (1764–1814). Karl Moritz Schumann wrote a monograph on Bignoniaceae in 1894 for Engler and Prantl's Die Natürlichen Pflanzenfamilien. After Schumann's monograph, no taxonomic treatment of the entire family was published until 2004. Classification In the APG IV system of classification for flowering plants, Bignoniaceae is one of the 24 families in the order Lamiales. (Lamiales has 25 families if Rehmanniaceae are accepted). Within the order, Bignoniaceae is in a group of eight families consisting of Thomandersiaceae, Pedaliaceae, Martyniaceae, Schlegeliaceae, Bignoniaceae, Verbenaceae, Acanthaceae, and Lentibulariaceae. This group is described as a polytomy, meaning no two of its members are known to be more closely related to each other than to any of the others. In the 20th century, the only issues of circumscription were whether Paulowniaceae and Schlegeliaceae should be merged into Bignoniaceae, or accepted as separate families. Crescentiina is composed of two strongly supported clades, informally named the Tabebuia alliance and the Paleotropical clade. The tribe Crescentieae is embedded in the Tabebuia alliance and might be expanded to include Spirotecoma. Perianthomega has been transferred from Tecomeae sensu stricto to Bignonieae, where it is sister to the remainder of the tribe. Roseodendron and Handroanthus were resurrected from Tabebuia in 2007. Mayodendron and Pachyptera have been resurrected. In 2009, Six of these groups have been recognized as tribes at one time or another, and are represented by their tribal names. Two of the groups are monogeneric and are designated by their constituent genera, Argylia and Delostoma. The other two groups are given informal names, pending a formal revision of the infrafamilial classification. Astianthus has never been sampled for DNA and its systematic position within the family remains obscure. Likewise, the placement of Romeroa in the Tabebuia alliance and the placement of Sphingiphila in Bignonieae are in doubt. Tecomaria is not included in the list below, and its recognition is controversial. It is monotypic (Tecomaria capensis), and had been long accepted, but was returned to Tecoma in 1980. A molecular phylogenetic study resolved it as sister to another South African genus, Podranea, but with only weak bootstrap support. Tecomaria has not yet been resurrected or transferred to another genus. The tribe Bignonieae has been the subject of considerable revision since 2006. Fischer et al. placed 46 genera in this tribe. Afterward, Perianthomega was transferred to it from Tecomeae sensu lato and Pachyptera was resurrected from Mansoa. Twenty-five of the genera of Fischer have been subsumed into other genera as follows: Gardnerodoxa into Neojobertia; Memora into Adenocalymma; Leucocalantha into Pachyptera; Pseudocatalpa, Paragonia, Periarrabidaea, Spathicalyx, and Ceratophytum into Tanaecium; Arrabidaea and Piriadacus into Fridericia; Clytostoma, Cydista, Macranthisiphon, Mussatia, Phryganocydia, Potamoganos, Roentgenia and Saritaea into Bignonia; also Distictis, Glaziovia, Haplolophium, and Pithecoctenium into Amphilophium. Thus, 23 genera are now recognized in Bignonieae. ;Incertae sedisAstianthus ;Tribe JacarandeaeJacaranda (synonym Digomphia) ;Tribe TourrettieaeEccremocarpusTourrettia ;Genus Argylia ;Tribe TecomeaeCampsidiumCampsisDeplancheaDinklageodoxaIncarvilleaLamiodendronNeosepicaeaPandoreaPodraneaTecomaTecomanthe ;Genus Delostoma ===Tribe Bignonieae=== • AdenocalymmaAmphilophium (synonym Distictella) • AnemopaegmaBignoniaCallichlamysCuspidariaDolichandraFridericiaLundiaManaosellaMansoaMartinellaNeojobertiaPachypteraPerianthomegaPleonotomaPyrostegiaStizophyllumTanaecium (synonym Sphingiphila) • TynanthusXylophragma ;Tribe OroxyleaeHierisMillingtoniaNyctocalosOroxylum ;Tribe CatalpeaeChilopsisCatalpa ===Crescentiina group=== ====Tabebuia alliance==== • AmphitecnaCrescentiaCybistaxEkmaniantheGodmaniaHandroanthusParatecomaParmentieraRomeroaRoseodendronSparattospermaSpirotecomaTabebuiaZeyheria Paleotropical cladeCatophractesColea (synonym Ophiocolea) • DolichandroneFernandoa (synonym Haplophragma) • HeterophragmaKigeliaMarkhamiaMayodendronNewbouldiaPajaneliaPauldopiaPerichlaenaPhyllarthronPhyllocteniumRadermacheraRhigozumRhodocoleaTecomellaSantisukiaSpathodeaStereospermum Obsolete generaArrabidaeaCeratophytumClytostomaCydistaDistictisGardnerodoxaGlazioviaHaplolophiumLeucocalanthaMacfadyenaMacranthisiphonMelloaMemoraMussatiaParabignoniaParagoniaPeriarrabidaeaPhryganocydiaPiriadacusPithecocteniumPotamoganosPseudocatalpaRoentgeniaSaritaeaSpathicalyx == Uses ==
Uses