Finch

The name Fringillidae for the finch family was introduced in 1819 by the English zoologist William Elford Leach in a guide to the contents of the British Museum. The taxonomy of the family, in particular the cardueline finches, has a long and complicated history. The study of the relationship between the taxa has been confounded by the recurrence of similar morphologies due to the convergence of species occupying similar niches. Beginning around 1990 a series of phylogenetic studies based on mitochondrial and nuclear DNA sequences resulted in substantial revisions in the taxonomy. Several groups of birds that had previously been assigned to other families were found to be related to the finches. The Neotropical Euphonia and the Chlorophonia were formerly placed in the tanager family Thraupidae due to their similar appearance but analysis of mitochondrial DNA sequences revealed that both genera were more closely related to the finches. They are now placed in a separate subfamily Euphoniinae within the Fringillidae. The Hawaiian honeycreepers were at one time placed in their own family, Drepanididae but were found to be closely related to the Carpodacus rosefinches and are now placed within the Carduelinae subfamily. The three largest genera, Carpodacus, Carduelis and Serinus were found to be polyphyletic. Each was split into monophyletic genera. The American rosefinches were moved from Carpodacus to Haemorhous. Carduelis was split by moving the greenfinches to Chloris and a large clade into Spinus leaving just three species in the original genus. Thirty seven species were moved from Serinus to Crithagra leaving eight species in the original genus. }} }} Fossil record Fossil remains of true finches are rare, and those that are known can mostly be assigned to extant genera at least. Like the other Passeroidea families, the true finches seem to be of roughly Middle Miocene origin, around 20 to 10 million years ago (Ma). An unidentifiable finch fossil from the Messinian age, around 12 to 7.3 million years ago (Ma) during the Late Miocene subepoch, has been found at Polgárdi in Hungary. ==Description==

The smallest "classical" true finches are the Andean siskin (Spinus spinescens) at as little as 9.5 cm (3.8 in) and the lesser goldfinch (Spinus psaltria) at as little as . The largest species is probably the collared grosbeak (Mycerobas affinis) at up to and , although larger lengths, to in the pine grosbeak (Pinicola enucleator), and weights, to in the evening grosbeak (Hesperiphona vespertina), have been recorded in species which are slightly smaller on average. They typically have strong, stubby beaks, which in some species can be quite large; however, Hawaiian honeycreepers are famous for the wide range of bill shapes and sizes brought about by adaptive radiation. All true finches have 9 primary remiges and 12 rectrices. The basic plumage colour is brownish, sometimes greenish; many have considerable amounts of black, while white plumage is generally absent except as wing-bars or other signalling marks. Bright yellow and red carotenoid pigments are commonplace in this family, and thus blue structural colours are rather rare, as the yellow pigments turn the blue color into green. Many, but by no means all true finches have strong sexual dichromatism, the females typically lacking the bright carotenoid markings of males. ==Distribution and habitat==

(Spinus tristis) male (left) and female (right) in Johnston County, North Carolina, USA The finches have a near-global distribution, being found across the Americas, Eurasia and Africa, as well as some island groups such as the Hawaiian islands. They are absent from Australasia, Antarctica, the Southern Pacific and the islands of the Indian Ocean, although some European species have been widely introduced in Australia and New Zealand. Finches are typically inhabitants of well-wooded areas, but some can be found on mountains or even in deserts. ==Behaviour==

eating coneflower seeds and taking flight, including slow motion. The finches are primarily granivorous, but euphoniines include considerable amounts of arthropods and berries in their diet, and Hawaiian honeycreepers evolved to utilize a wide range of food sources, including nectar. The diet of Fringillidae nestlings includes a varying amount of small arthropods. True finches have a bouncing flight like most small passerines, alternating bouts of flapping with gliding on closed wings. Most sing well and several are commonly seen cagebirds; foremost among these is the domesticated canary (Serinus canaria domestica). The nests are basket-shaped and usually built in trees, more rarely in bushes, between rocks or on similar substrate. ==List of genera==

The family Fringillidae contains 235 species divided into 50 genera and three subfamilies. The subfamily Carduelinae includes 18 extinct Hawaiian honeycreepers and the extinct Bonin grosbeak. See List of Fringillidae species for further details. Subfamily Fringillinae • Fringilla – 5 species of chaffinch, 2 species of blue chaffinch, and the brambling Subfamily Carduelinae • Mycerobas – 4 Palearctic grosbeaks • Coccothraustes – 3 species • Eophona – 2 oriental grosbeaks, the Chinese and the Japanese grosbeak • Pinicola – pine grosbeak • Pyrrhula – 8 bullfinch species • Rhodopechys – 2 species, the Asian crimson-winged finch and the African crimson-winged finch • Bucanetes – trumpeter and the Mongolian finch • Agraphospiza – Blanford's rosefinch • Callacanthis – spectacled finch • Pyrrhoplectes – golden-naped finch • Procarduelis – dark-breasted rosefinch • Leucosticte – 6 species of mountain and rosy finches • Carpodacus – 28 Palearctic rosefinch species • Hawaiian honeycreeper group (tribe Drepanidini) • Melamprosops – contains a single extinct species, the po'ouli • Paroreomyza – 3 species, the Oahu alauahio, the Maui alauahio and the extinct kakawahie • Oreomystis – akikiki • Telespiza – 4 species, the Laysan finch, the Nihoa finch, and 2 prehistoric species • Loxioides – 2 species, the palila and a prehistoric species • Rhodacanthis – 2 recently extinct species, the lesser and the greater koa finch, and 2 prehistoric species • Chloridops – extinct species, the Kona grosbeak • Psittirostra – ou • Dysmorodrepanis – extinct species, the Lanai hookbill • Drepanis – 2 extinct species, the Hawaii mamo and the black mamo, and the extant iiwi • Ciridops – single recently extinct species, the Ula-ai-hawane, and 3 prehistoric species • Palmeria – contains a single species, the akohekohe • Himatione – 2 species, the apapane and the extinct Laysan honeycreeper • Viridonia – single extinct species, the greater amakihi • Akialoa – 4 recently extinct species, and 2 prehistoric species • Hemignathus – 4 species, only one of which is extant • Pseudonestor – Maui parrotbill • Magumma – anianiau • Loxops – 5 species, of which one is extinct • Chlorodrepanis – 3 species, the Hawaii, Oahu and Kauai amakihi • Haemorhous – 3 North America rosefinches • Chloris – 6 greenfinches • Rhodospiza – desert finch • Rhynchostruthus – 3 golden-winged grosbeaks • Linurgus – oriole finch • Crithagra – 37 species of canaries, serins and siskins from Africa and the Arabian Peninsula • Linaria – 4 species including the twite and three linnets • Acanthis – 3 redpolls • Loxia – 6 crossbills • Chrysocorythus – 2 species • Carduelis – 3 species including the European goldfinch • Serinus – 8 species including the European serin • Spinus – 20 species including the North American goldfinches and the Eurasian siskin Subfamily Euphoniinae • Euphonia – 27 species all with euphonia in their English name • Chlorophonia – 5 species all with chlorophonia in their English name ==Gallery==

File:Coccothraustes coccothraustes 1 (Marek Szczepanek).jpg|Hawfinch (Coccothraustes coccothraustes), one of the Holarctic grosbeaks File:Cassin's Finch (male).jpg|Cassin's finch (Haemorhous cassinii), an American rosefinch File:Carpodacus roseus.jpg|Pallas' rosefinch (Carpodacus roseus), a true rosefinch File:PINTASSILGO ( Carduelis magellanica ).jpg|Hooded siskin (Spinus magellanicus) File:Vestiaria coccinea -Hawaii -adult-8 (4).jpg|ʻIʻiwi (Drepanis coccinea), a Hawaiian honeycreeper File:Euphonia violacea-2.jpg|Male violaceous euphonia (Euphonia violacea) File:Carduelis carduelis close up.jpg|European goldfinch (Carduelis carduelis) File:Chloris chloris.jpg|European greenfinch (Chloris chloris) File:Pinzón azul de Gran Canaria (macho), M. A. Peña.jpg|Gran Canaria blue chaffinch (Fringilla polatzeki) File:Teidefink.jpg|Tenerife blue chaffinch (Fringilla teydea) File:Euphonia elegantissima.jpg|Elegant euphonia (Chlorophonia elegantissima) File:Desert Finch - Uzbekistan S4E7626-2 (23039952955).jpg|Desert finch (Rhodospiza obsoleta) File:Pine Grosbeak (Pinicola enucleator) (13667564073).jpg|Pine grosbeak (Pinicola enucleator) File:Hesperiphona vespertina CT3.jpg|Evening grosbeak (Hesperiphona vespertina) File:Chloris spinoides.jpg|Yellow-breasted greenfinch (Chloris spinoides) File:Serinus flaviventris 2013 03 09.jpg|Yellow canary (Crithagra flaviventris) File:Streaky Seedeater, Ngorongoro Crater (8495906768).jpg|Streaky seedeater (Crithagra striolata) File:Fringilla coelebs moreletti.png|Azores chaffinch (Fringilla coelebs moreletti) ==See also==