From the
Ediacaran (600 ) into the
Devonian (360 ), the
Proto-Tethys Ocean existed and was situated between
Baltica and
Laurentia to the north and
Gondwana to the south. From the
Silurian (440 ) through the
Jurassic periods, the Paleo-Tethys Ocean existed between the
Hunic terranes and Gondwana. Over a period of 400 million years, continental
terranes intermittently separated from Gondwana in the Southern Hemisphere to migrate northward to form Asia in the Northern Hemisphere.
Triassic Period About 250 Mya, during the
Triassic, a new ocean began forming in the southern end of the Paleo-Tethys Ocean. A rift formed along the northern continental shelf of Southern
Pangaea (Gondwana). Over the next 60 million years, that piece of shelf, known as
Cimmeria, traveled north, pushing the floor of the Paleo-Tethys Ocean under the eastern end of northern
Pangaea (early / proto-
Laurasia). The Neo-Tethys Ocean formed between Cimmeria and Gondwana, directly over where the Paleo-Tethys formerly rested.
Jurassic Period During the
Jurassic period about 150 Mya, Cimmeria finally collided with Laurasia and stalled, so the ocean floor behind it
buckled under, forming the
Tethys Trench. Water levels rose, and the western Tethys shallowly covered significant portions of Europe, forming the first Tethys Sea. Around the same time, Laurasia and Gondwana began
drifting apart, opening an extension of the Tethys Sea between them, which today is the part of the Atlantic Ocean between the
Mediterranean and the
Caribbean. As North and South America were still attached to the rest of Laurasia and Gondwana, respectively, the Tethys Ocean in its widest extension was part of a continuous oceanic belt running around the Earth between about
latitude 30°N and the
Equator. Thus,
ocean currents at the time around the
Early Cretaceous ran very differently from the way they do today.
Late Cretaceous Between the Jurassic and the
Late Cretaceous, which started about 100 Mya, Gondwana began breaking up, pushing Africa and India north across the Tethys and opening up the Indian Ocean. During the Late Cretaceous, the Tethys Sea was home to many different animals, including
marine reptiles,
bony fish,
cartilaginous fish, and
cephalopods. The
islands that were located in the northern parts of the Tethys Sea (
Europe) created
biodiverse ecosystems that had animals that went through
insular dwarfism and
insular gigantism. The insular dwarfism process happened mostly to the
dinosaurs that lived on the islands, like the
sauropods and the
hadrosaurs.
Telmatosaurus is a good representation of the insular dwarfism process. While the insular dwarfism process happened to the dinosaurs, the
pterosaurs that lived on the islands went through the process known as insular gigantism.
Hatzegopteryx was a huge
azhdarchid pterosaur that lived on the islands of the Tethys Sea. This giant pterosaur would have filled its
ecological niche as an
apex predator. During the
Maastrichtian, the Tethys Sea had many different large mosasaurs that lived in the same geographical area and would have competed with each other. Europe had large mosasaurs like
Prognathodon giganteus, Prognathodon saturator, Prognathodon sectorius,
Mosasaurus hoffmannii, and Mosasaurus lemonnieri.
North Africa would have also had large mosasaurs like Prognathodon giganteus, Prognathodon currii,
Thalassotitan atrox,
Hainosaurus boubker, and Mosasaurus beaugei. The competition between many different apex predators is something we don't only see in the Tethys Sea, but also in the
Western Interior Seaway.
Cenozoic (top image), but by the Oligocene, most of this had dried out (bottom image), and the Tethys was almost entirely divided into the Indian Ocean, Mediterranean and Paratethys. Throughout the
Cenozoic (66 million years to the dawn of the Neogene, 23 Mya), the connections between the Atlantic and Indian Oceans across the Tethys were eventually closed off in what is now the Middle East during the
Miocene, as a consequence of the northern migration of Africa/Arabia and global sea levels falling due to the concurrent formation of the
Antarctic Ice Sheet. This decoupling occurred in two steps, first around 20 Mya and another around 14 Mya. Separation from the wider Tethys during the early Miocene initially led to a boost in
primary productivity for the Paratethys, but this gave way to a total ecosystem collapse during the late Miocene as a result of rapid dissolution of
carbonate. ==Historical theory==