The Margaret Formation comprises
sandstones, sandy
siltstones, clay-rich sandstones, coal seams and clay-rich
coal seams and volcanic ash beds. The thickness of the formation, which overlies the
Mount Moore Formation, reaches about .
Dating A volcanic ash layer containing
crandallite in the middle of the formation was dated using U-Pb radiometric dating in 2017 to 53.7 ± 0.6 Ma. In 2010, ashes of the formation were dated to 52.6 ± 1.9 Ma.
Arctic climate of the Early Eocene During the Early Eocene, the climate of much of northern North America was warm and wet, with mean annual temperatures (MAT) as high as , mean annual precipitation (MAP) of , mild frost-free winters (coldest month mean temperature >), and climatic conditions that supported extensive temperate forest ecosystems. Ensemble estimates of mean annual temperatures for the high-latitude fossil localities in Arctic Canada ranged from , with the range of coldest month mean temperature from and warmest month mean temperatures from . Mean annual precipitation estimates for the Margaret Formation ranged between . The mean summer precipitation has been estimated at and mean winter precipitation at . The fossils and sedimentology indicate a lush, rain forest community on a coastal delta plain. Multiple palaeoclimate proxies, ranging from oxygen isotope analysis of vertebrate bones and teeth to palaeofloral analyses, estimate a mild temperate climate for the Eocene High Arctic, where winters remained at or just above freezing and summer temperatures extended to or higher. These temperatures are a far cry from today's High Arctic, where central Ellesmere Island experiences a mean annual temperature of , a warm month mean temperature of about and a cold month mean temperature of or colder. Despite the mild Eocene Arctic climate, the vertebrate fauna would have experienced months of total darkness and cooler temperatures during the winter. Recent isotopic work suggests that some mammals, including the hippo-like
Coryphodon, were year-round residents in the High Arctic. Given that
Gastornis was large (approaching ) and flightless, it likely also was a year-round resident of the Arctic. In contrast, the volant
Presbyornis might have been a seasonal migrant to the Arctic. == Fossil content ==