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Orange (fruit)

The orange, also called sweet orange to distinguish it from the bitter orange, is the fruit of a tree in the family Rutaceae. Botanically, this is the hybrid Citrus × sinensis, between the pomelo and the mandarin orange. The chloroplast genome, and therefore the maternal line, is that of pomelo. Hybrids of the sweet orange form later types of mandarin and the grapefruit. The sweet orange has had its full genome sequenced.

Description
The orange tree is a relatively small evergreen, flowering tree, with an average height of , although some very old specimens can reach . Its oval leaves, which are alternately arranged, are long and have crenulate margins. Sweet oranges grow in a range of different sizes, and shapes varying from spherical to oblong. Inside and attached to the rind is a porous white tissue, the white, bitter mesocarp or albedo (pith). The orange contains a number of distinct carpels (segments or pigs, botanically the fruits) inside, typically about ten, each delimited by a membrane and containing many juice-filled vesicles and usually a few pips. When unripe, the fruit is green. The grainy irregular rind of the ripe fruit can range from bright orange to yellow-orange, but frequently retains green patches or, under warm climate conditions, remains entirely green. Like all other citrus fruits, the sweet orange is non-climacteric, not ripening off the tree. The Citrus sinensis group is subdivided into four classes with distinct characteristics: common oranges, blood or pigmented oranges, navel oranges, and acidless oranges. File:Orange Blossom.JPG|Flowers File:Orange tree fruiting.jpg|Fruit starting to develop File:OrangeBloss wb.jpg|Flowers and fruit simultaneously File:Laranxeira Naranjo GFDL.JPG|Mature tree in Galicia, Spain, fruiting in November File:Structure of an orange.svg|Structure of the botanical hesperidium == History ==
History
Hybrid origins Citrus trees are angiosperms, and most species are almost entirely interfertile. This includes grapefruits, lemons, limes, oranges, and many citrus hybrids. As the interfertility of oranges and other citrus has produced numerous hybrids and cultivars, and bud mutations have also been selected, citrus taxonomy has proven difficult. The sweet orange, Citrus x sinensis, is not a wild fruit, but arose in domestication in East Asia. It originated in a region encompassing Southern China, Northeast India, and Myanmar. The fruit was created as a cross between a non-pure mandarin orange and a hybrid pomelo that had a substantial mandarin component. Since its chloroplast DNA is that of pomelo, it was likely the hybrid pomelo, perhaps a pomelo BC1 backcross, that was the maternal parent of the first orange. Based on genomic analysis, the relative proportions of the ancestral species in the sweet orange are approximately 42% pomelo and 58% mandarin. All varieties of the sweet orange descend from this prototype cross, differing only by mutations selected for during agricultural propagation. The citranges are a group of sweet orange x trifoliate orange (Citrus trifoliata) hybrids. of mandarin and pomelo. Large-scale cultivation started in the 10th century, as evidenced by complex irrigation techniques specifically adapted to support orange orchards. At Versailles, potted orange trees in solid silver tubs were placed throughout the rooms of the palace, while the Orangerie allowed year-round cultivation of the fruit to supply the court. When Louis condemned his finance minister, Nicolas Fouquet, in 1664, part of the treasures that he confiscated were over 1,000 orange trees from Fouquet's estate at Vaux-le-Vicomte. To the Americas Spanish travelers introduced the sweet orange to the American continent. On his second voyage in 1493, Christopher Columbus may have planted the fruit on Hispaniola. Subsequent expeditions in the mid-1500s brought sweet oranges to South America and Mexico, and to Florida in 1565, when Pedro Menéndez de Avilés founded St Augustine. Spanish missionaries brought orange trees to Arizona between 1707 and 1710, while the Franciscans did the same in San Diego, California, in 1769. Florida farmers obtained seeds from New Orleans around 1872, after which orange groves were established by grafting the sweet orange on to sour orange rootstocks. The 1870s saw the introduction of improved fruit varieties. In 1873, navel orange plants from Brazil were distributed by the U.S. Department of Agriculture. Luther C. Tibbets and Eliza Tibbets successfully cultivated these in Riverside, leading to widespread planting of the sweet, seedless navel orange, which became the backbone of the California citrus industry. The Valencia orange, introduced in 1876, matured in summer and fall, complementing the winter-ripening navel and providing oranges year-round. The completion of major railroads (Southern Pacific in 1877, and the Santa Fe in 1885) and the introduction of ventilated boxcars revolutionized distribution, opening national markets and triggering a planting frenzy in southern California. By 1885, the number of citrus trees in California had grown from 90,000 (in 1875) to 2 million, and to 4.5 million by 1901. The 1890s brought pest control advances (spraying, fumigation) and frost protection (heaters, later wind machines). The University of California established its Citrus Experiment Station in 1907, supporting research and innovation. Cooperative marketing emerged with the formation of the California Fruit Growers Exchange in 1905, later known as Sunkist Growers Inc., which helped standardize and market California citrus worldwide. By the 1980s, California was the second largest orange producer in the U.S., after Florida. == Etymology ==
Etymology
The word "orange" has its etymological roots in the Dravidian language family of South India. From there, the word passed to Sanskrit (), meaning 'orange tree'. The Sanskrit word reached European languages through Persian () and its Arabic derivative (). The word entered Late Middle English in the 14th century via Old French . Other forms include Old Provençal , Italian arancia, formerly narancia. with the first recorded use of orange as a color name in English in 1512. == Composition ==
Composition
Nutrition Orange flesh is 87% water, 12% carbohydrates, 1% protein, and contains negligible fat (table). In a reference amount of , orange flesh provides 47 calories, and is a rich source of vitamin C, providing 59% of the Daily Value, with no other micronutrients in significant amounts (table). Phytochemicals Oranges contain diverse phytochemicals, including carotenoids (beta-carotene, lutein and beta-cryptoxanthin), flavonoids (e.g. naringenin) and numerous volatile organic compounds producing orange aroma, including aldehydes, esters, terpenes, alcohols, and ketones. Orange juice contains only about one-fifth the citric acid of lime or lemon juice (which contain about 47 g/L). Taste , a volatile compound contributing to the fragrance of oranges The taste of oranges is determined mainly by the ratio of sugars to acids, whereas orange aroma derives from volatile organic compounds, including alcohols, aldehydes, ketones, terpenes, and esters. Bitter limonoid compounds, such as limonin, decrease gradually during development, whereas volatile aroma compounds tend to peak in mid- to late-season development. Taste quality tends to improve later in harvests when there is a higher sugar/acid ratio with less bitterness. to 4.0. Taste and aroma vary according to genetic background, environmental conditions during development, ripeness at harvest, postharvest conditions, and storage duration. == Cultivars ==
Cultivars
Common Common oranges (also called "white", "round", or "blond" oranges) constitute about two-thirds of all orange production. The majority of this crop is used for juice. Navel Navel oranges have a characteristic second fruit at the apex, which protrudes slightly like a human navel. They are mainly an eating fruit, as their thicker skin makes them easy to peel, they are less juicy and their bitterness makes them less suitable for juice. The cultivar rapidly spread to other countries, but being seedless it had to be propagated by cutting and grafting. The Cara Cara is a type of navel orange grown mainly in Venezuela, South Africa and California's San Joaquin Valley. It is sweet and low in acid, with distinctively pinkish red flesh. It was discovered at the Hacienda Cara Cara in Valencia, Venezuela, in 1976. Blood Blood oranges, with an intense red coloration inside, are widely grown around the Mediterranean; there are several cultivars. The redness is mainly due to the anthocyanin pigment chrysanthemin (cyanidin 3-O-glucoside). Acidless Acidless oranges are an early-season fruit with very low levels of acid. They also are called "sweet" oranges in the United States, with similar names in other countries: douce in France, sucrena in Spain, dolce or maltese in Italy, meski in North Africa and the Near East (where they are especially popular), succari in Egypt, and lima in Brazil. The lack of acid, which protects orange juice against spoilage in other groups, renders them generally unfit for processing as juice, so they are primarily eaten. They remain profitable in areas of local consumption, but rapid spoilage renders them unsuitable for export to major population centres of Europe, Asia, or the United States. File:Florida orange grove.JPG|A grove of Valencia oranges in Florida File:Cara cara orange cut in half.JPG|Cara Cara navel orange File:BloodOrange.jpg|Blood orange == Cultivation ==
Cultivation
Climate Like most citrus plants, oranges do well under moderate temperatures—between —and require considerable amounts of sunshine and water. They are principally grown in tropical and subtropical regions. Another procedure involves burning fuel oil in smudge pots put between the trees. These burn with a great deal of particulate emission, so condensation of water vapor on the particulate soot prevents condensation on plants and raises the air temperature very slightly. Smudge pots were developed after a disastrous freeze in southern California in January 1913 destroyed a whole crop. Propagation Commercially grown orange trees are propagated asexually by grafting a mature cultivar onto a suitable seedling rootstock to ensure the same yield, identical fruit characteristics, and resistance to diseases throughout the years. Propagation involves two stages: first, a rootstock is grown from seed. Then, when it is approximately one year old, the leafy top is cut off and a bud taken from a specific scion variety, is grafted into its bark. The scion is what determines the variety of orange, while the rootstock makes the tree resistant to pests and diseases and adaptable to specific soil and climatic conditions. Thus, rootstocks influence the rate of growth and have an effect on fruit yield and quality. Rootstocks must be compatible with the variety inserted into them because otherwise, the tree may decline, be less productive, or die. and that farmers can combine the best attributes of a scion with those of a rootstock. Harvest Canopy-shaking mechanical harvesters are being used increasingly in Florida to harvest oranges. Current canopy shaker machines use a series of six-to-seven-foot-long tines to shake the tree canopy at a relatively constant stroke and frequency. Oranges are picked once they are pale orange. Degreening Oranges must be mature when harvested. In the United States, laws forbid harvesting immature fruit for human consumption in Texas, Arizona, California, and Florida. Ripe oranges, however, often have some green or yellow-green color in the skin. Ethylene gas is used to turn green skin to orange. This process is known as "degreening", "gassing", "sweating", or "curing". Storage Commercially, oranges can be stored by refrigeration in controlled-atmosphere chambers for up to twelve weeks after harvest. Storage life ultimately depends on cultivar, maturity, pre-harvest conditions, and handling. At home, oranges have a shelf life of about one month, and are best stored loose. File:CSIRO ScienceImage 4314 Spraying oranges in an orchard at Griffith NSW 2002.jpg|Spraying oranges in an orchard in Australia File:California Orange Grove2.jpg|Orange grove in California 100521 picking oranges in moshav zimrat PikiWiki Israel.jpg|Picking oranges, Israel File:100535 picking oranges in moshav zimrat PikiWiki Israel.jpg|Harvest, Israel File:2010-12-14 Maroc Agadir Soukh local market.jpg|Market stall, Morocco == Pests and diseases ==
Pests and diseases
Pests insects devastated orange groves across California in the 19th century, and were the first pest to be subject to successful biological control. Diseases '', is a major vector of citrus greening disease. As from 2009, 0.87% of the trees in Brazil's main orange growing areas (São Paulo and Minas Gerais) showed symptoms of greening, an increase of 49% over 2008. The disease is spread primarily by psyllid plant lice such as the Asian citrus psyllid (Diaphorina citri Kuwayama), an efficient vector of the bacterium. Foliar insecticides reduce psyllid populations for a short time, but also suppress beneficial predatory ladybird beetles. Soil application of aldicarb provided limited control of Asian citrus psyllid, while drenches of imidacloprid to young trees were effective for two months or more. Management of citrus greening disease requires an integrated approach that includes use of clean stock, elimination of inoculum via voluntary and regulatory means, use of pesticides to control psyllid vectors in the citrus crop, and biological control of the vectors in non-crop reservoirs. == Production ==
Production
In 2022, world production of oranges was 76 million tonnes, led by Brazil with 22% of the total, followed by India, China, and Mexico. In the United States, groves are located mainly in Florida, California, and Texas. The majority of California's crop is sold as fresh fruit, whereas Florida's oranges are destined to juice products. The Indian River area of Florida produces high quality juice, which is often sold fresh and blended with juice from other regions, because Indian River trees yield sweet oranges but in relatively small quantities. == Culinary use ==
Culinary use
Dessert fruit and juice Oranges, whose flavor may vary from sweet to sour, are commonly peeled and eaten fresh raw as a dessert. Orange juice is obtained by squeezing the fruit on a special tool (a juicer or squeezer) and collecting the juice in a tray or tank underneath. This can be made at home or, on a much larger scale, industrially. Orange juice is a traded commodity on the Intercontinental Exchange. Frozen orange juice concentrate is made from freshly squeezed and filtered juice. Marmalade Oranges are made into jam in many countries; in Britain, bitter Seville oranges are used to make marmalade. Almost the whole Spanish production is exported to Britain for this purpose. The entire fruit is cut up and boiled with sugar; the pith contributes pectin, which helps the marmalade to set. The first recipe was by an Englishwoman, Mary Kettilby, in 1714. Pieces of peel were first added by Janet Keiller of Dundee in the 1790s, contributing a distinctively bitter taste. Orange peel contains the bitter substances limonene and naringin. Extracts Zest is scraped from the coloured outer part of the peel, and used as a flavoring and garnish in desserts and cocktails. Sweet orange oil is a by-product of the juice industry produced by pressing the peel. It is used for flavoring food and drinks; it is employed in the perfume industry and in aromatherapy for its fragrance. The oil consists of approximately 90% D-limonene, a solvent used in household chemicals such as wood conditioners for furniture and—along with other citrus oils—detergents and hand cleansers. It is an efficient cleaning agent with a pleasant smell, promoted for being environmentally friendly and therefore preferable to petrochemicals. It is, however, irritating to the skin and toxic to aquatic life. File:Oranges and orange juice.jpg|Fruit and juice File:Zesting an orange.jpg|Zesting an orange File:Sevilleorangemarmalade.jpg|Homemade marmalade, England == In human culture ==
In human culture
Oranges have featured in human culture since ancient times. The earliest mention of the sweet orange in Chinese literature dates from 314 BC. Pham comments that the Arnolfini Portrait by Jan van Eyck contains in a small detail one of the first representations of oranges in Western art, the costly fruit perhaps traded by the merchant Arnolfini himself. By the 17th century, orangeries were added to great houses in Europe, both to enable the fruit to be grown locally and for prestige, as seen in the Versailles Orangerie completed in 1686. The Dutch Post-Impressionist artist Vincent van Gogh portrayed oranges in paintings such as his 1889 Still Life of Oranges and Lemons with Blue Gloves and his 1890 A Child with Orange, both works late in his life. The American artist of the Ashcan School, John Sloan, made a 1935 painting Blond Nude with Orange, Blue Couch, while Henri Matisse's last painting was his 1951 Nude with Oranges; after that he only made cut-outs. File:Yellow Oranges and Green Tangerines (橙黃橘綠) by Zhao Lingrang (趙令穰).jpg|Yellow Oranges and Green Tangerines by Zhao Lingrang, Chinese fan painting from the Song dynasty, c. 1070–1100 File:Jan van Eyck 002.jpg|Detail of the Arnolfini Portrait by Jan van Eyck, 1434 File:Primavera (Botticelli) (detail).jpg|Detail of Primavera by Sandro Botticelli, 1482, set in an orange grove File:Måleri, stilleben. Frukt - Skoklosters slott - 88970.tif|Still life with oranges on a plate. Possibly Jacques Linard or Louise Moillon, 1640 File:Orangerie du château de Versailles le 11 septembre 2015 - 78.jpg|The Versailles Orangerie, 1686 File:OUDRY Orange Tree.jpg|Jean-Baptiste Oudry, The Orange Tree, 1740 File:Vincent van Gogh, Still Life of Oranges and Lemons with Blue Gloves, 1889, NGA 164923.jpg| Still Life of Oranges and Lemons with Blue Gloves by Vincent van Gogh, 1889 == See also ==
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