Sloth bear

George Shaw in 1791 named the species Bradypus ursinus. In 1793, Meyer named it Melursus lybius, and in 1817, de Blainville named it Ursus labiatus because of its long lips. Illiger named it Prochilus hirsutus, the Greek genus name indicating long lips, while the specific name noted its long and coarse hair. Fischer called it Chondrorhynchus hirsutus, while Tiedemann named it Ursus longirostris. Subspecies and range == Evolution ==

Sloth bears may have reached their current form in the Early Pleistocene, the time when the bear family specialised and dispersed. A fragment of fossilised humerus from the Pleistocene, found in Andhra Pradesh's Kurnool Basin is identical to the humerus of a modern sloth bear. The fossilised skulls of a bear once named Melursus theobaldi found in the Shivaliks from the Early Pleistocene or Early Pliocene are thought by certain authors to represent an intermediate stage between sloth bears and ancestral brown bears. M. theobaldi itself had teeth intermediate in size between sloth bears and other bear species, though its palate was the same size as the former species, leading to the theory that it is the sloth bear's direct ancestor. Sloth bears probably arose during the Middle Pliocene and evolved in the Indian subcontinent. The sloth bear shows evidence of having undergone a convergent evolution similar to that of other ant-eating mammals. The sloth bear is one of eight extant species in the bear family Ursidae and of six extant species in the subfamily Ursinae. == Characteristics ==

Sloth bears adults are medium-sized bears. The typical weight range for females is from , and for males is from . Exceptionally large female specimens can reach and males up to . The average weight of sloth bears from the nominate subspecies in Nepal was in females and in males. Nominate bears in India were found to weigh average in males and in female per one study. Specimens from Sri Lanka (M. u. inornatus) may weigh up to in females and in males. However six Sri Lankan male sloth bears averaged only , and was the average for four females, so Sri Lankan bears could be around 30% lighter in body mass than nominate race bears and with more pronounced size sexual dimorphism. They are high at the shoulder, and have a body length of . Besides being smaller than males, females reportedly typically have more fur between their shoulders. Sloth bear muzzles are thick and long, with small jaws and bulbous snouts with wide nostrils. They have long lower lips which can be stretched over the outer edge of their noses, and they lack upper incisors, thus allowing them to suck up large numbers of insects. The premolars and molars are smaller than in other bears, as they do not chew as much vegetation. In adults, the teeth are usually in poor condition, due to the amount of soil they suck up and chew when feeding on insects. This feature, which is also present in Asian black bears and sun bears, is thought to serve as a threat display, as all three species are sympatric with tigers (tigers usually do not carry out attacks on an adult bear if the bear is aware or facing the cat). == Distribution and habitat ==

The sloth bear's global range includes India, the Terai of Nepal, temperate climatic zones of Bhutan and Sri Lanka. It occurs in a wide range of habitats including moist and dry tropical forests, savannahs, scrublands and grasslands below on the Indian subcontinent, and below in Sri Lanka's dry forests. It is regionally extinct in Bangladesh. == Behaviour and ecology ==

Adult sloth bears may travel in pairs. Males are often observed to be gentle with cubs. They may fight for food. They walk in a slow, shambling motion, with their feet being set down in a noisy, flapping motion. They are capable of galloping faster than running humans. Although they appear slow and clumsy, both young and adult sloth bears are excellent climbers. They occasionally will climb to feed and to rest, though not to escape enemies, as they prefer to stand their ground. Sloth bear mothers carry their cubs up trees as the primary defense against attacks by predators instead of sending them up trees. The cubs can be threatened by predators such as tigers, leopards, and other bears. They are adequate climbers on more accessible trees but cannot climb as quickly or on as varied surfaces as can black bears due to the sloth species' more elongated claw structure. Given their smaller size and still shorter claws, sloth bear cubs probably climb more proficiently than adults (much as brown bear cubs can climb well but not adults). Reproduction , India The breeding season for sloth bears varies according to location: in India, they mate in April, May, and June, and give birth in December and early January, while in Sri Lanka, it occurs all year. Sows gestate for 210 days, and typically give birth in caves or in shelters under boulders. Litters usually consist of one or two cubs, or rarely three. On arriving at a mound, they scrape at the structure with their claws until they reach the large combs at the bottom of the galleries, and disperse the soil with violent puffs. The termites are then sucked up through the muzzle, producing a sucking sound which can be heard 180 m away. Rarely, sloth bears can become addicted to sweets in hotel waste, visiting rubbish bins, even inside populated towns, all year long. Their diet includes animal flesh. In Neyyar Wildlife Sanctuary, Kerala, seeds of six tree species eaten and excreted by sloth bears (Artocarpus hirsuta, A. integrifolia, Cassia fistula, Mangifera indica, Zizyphus oenoplina) did not see significantly different percentages of germination (appearance of cotyledon) when compared to germinated seeds that had not been passed through the gut of the bears. However, seeds germinated much faster after being ingested by bears for three species, Artocarpus hirsuta, Cassia fistula, and Zizyphus oenoplina. This experiment suggests that sloth bears may play an important role in seed dispersal and germination, with effects varying by tree species. Bengal tigers occasionally prey on sloth bears. Tigers usually give sloth bears a wide berth, though some specimens may become habitual bear killers, and it is not uncommon to find sloth bear fur in tiger scats. Tigers typically hunt sloth bears by waiting for them near termite mounds, then creeping behind them and seizing them by the back of their necks and forcing them to the ground with their weight. As tigers are known to mimic the calls of sambar deer to attract them, sloth bears react fearfully even to the sounds made by deer themselves. In 2011, a female bear with cubs was observed to stand her ground and prevail in a confrontation against two tigers (one female, one male) in rapid succession. Besides tigers there are few predators of sloth bears. Leopards can also be a threat, as they are able to follow sloth bears up trees. Sloth bears occasionally chase leopards from their kills. When attacking them, dholes try to prevent the bear from retreating into caves. Unlike tigers which prey on sloth bears of all size, there is little evidence that dholes are a threat to fully-grown sloth bears other than exceptionally rare cases. In one case, a golden jackal (a species much smaller and less powerful than a sloth bear and not generally a pack hunter as is the dhole) was seen to aggressively displace an adult bear which passively loped away from the snapping canid, indicating the sloth bear does not regard other carnivores as competition. Sloth bears are sympatric with Asiatic black bears in northern India, and the two species, along with the sun bear, coexist in some of the national parks and wildlife sanctuaries. They are also found together in Assam, Manipur, and Mizoram, in the hills south of the Brahmaputra River, the only places occupied by all three bear species. The three species do not act aggressively toward each other. This may be because the three species generally differ in habit and dietary preferences. Asian elephants apparently do not tolerate sloth bears in their vicinity. The reason for this is unknown, as individual elephants known to maintain their composure near tigers have been reported to charge bears. The Indian rhinoceros has a similar intolerance for sloth bears, and will charge at them. == Status and conservation ==

The sloth bear is listed in Schedule I of the Indian Wildlife Protection Act, 1972, which provides for its legal protection. Commercial international trade of the sloth bear, including parts and derivatives, is prohibited as it is listed in Appendix I of the Convention on International Trade in Endangered Species. The population of sloth bears grows when they live in high-profile reserves that protect species, such as tigers and elephants. Directly managed reserves could conserve the sloth bear, hence such reserves must be supported. Managing garbage, especially hotel waste with foods, is essential in situations where sloth bears get used to entering towns with an increase in the number of accidental attacks on humans. Major sloth bear sanctuaries in India include the Daroji Sloth Bear Sanctuary in Karnataka, and Jessore Sloth Bear Sanctuary in Gujarat. A Sloth Bear Conservation Reserve is proposed in Mirzapur district of Uttar Pradesh. == Relationships with humans ==

Attacks on humans , Dahod district, Gujarat, India Sloth bears are one of the most aggressive extant bears and, due to large human populations often closely surrounding reserves that hold bears, aggressive encounters and attacks are relatively frequent, though in some places, attacks appear to be a reaction to encountering people accidentally. Sloth bears likely view humans as potential predators, as their reactions to them (roaring, followed by retreat or charging) are similar to those evoked in the presence of tigers and leopards. According to Robert Armitage Sterndale, in his Mammalia of India (1884, p. 62): Captain Williamson in his Oriental Field Sports wrote of how sloth bears rarely killed their human victims outright, but would suck and chew on their limbs until they were reduced to bloody pulps. One specimen, known as the sloth bear of Mysore, was responsible for the deaths of 12 people and the mutilation of 24 others. It was shot by Kenneth Anderson. Although sloth bears have attacked humans, they rarely become man-eaters. Dunbar-Brander's Wild Animals of Central India mentions a case in which a sow with two cubs began a six-week reign of terror in Chanda, a district of the Central Provinces, during which more than one of their victims had been eaten, while the sloth bear of Mysore partially ate at least three of its victims. Unlike some other bear species, which at times make mock charges at humans when surprised or frightened without making physical contact, sloth bears frequently appear to initiate a physical attack almost immediately. When people living near an aggressive population of sloth bears were armed with rifles, it was found that it was an ineffective form of defense, since the bear apparently charges and knocks the victim back (often knocking the rifle away) before the human has the chance to defend themself. In Madhya Pradesh, sloth bear attacks accounted for the deaths of 48 people and the injuring of 686 others between 1989 and 1994, probably due in part to the density of population and competition for food sources. A total of 137 attacks (resulting in 11 deaths) occurred between April 1998 and December 2000 in the North Bilaspur Forest Division of Chhattisgarh. The majority of attacks were perpetrated by single bears, and occurred in kitchen gardens, crop fields, and in adjoining forests during the monsoon season. One Mr. Watts Jones wrote a first-hand account of how it feels to be attacked by a sloth bear, recalling when he failed to score a direct hit against a bear he had targeted: In 2016, according to a forest official, a female bear had killed three people, and hurt five others in Gujarat State's Banaskantha district, near Balaram Ambaji Wildlife Sanctuary, with some of the casualties being colleagues. At first, an attempt was made to trace and cage it, but this failed, costing the life of one official, and so a team of both officials and policemen shot the bear. In Mount Abu town in southern Rajasthan, sloth bears attacked people inside towns where they were seeking hotel waste in rubbish bins and encountered people by chance. Sloth bears were also occasionally speared on horseback. Dancing bears were historically a popular entertainment in India, dating back to the 13th century and the pre-Mughal era. The Kalandars, who practised the tradition of capturing sloth bears for entertainment purposes, were often employed in the courts of Mughal emperors to stage spectacles involving trained bears. In 2009, following a seven-year campaign by a coalition of Indian and international animal welfare groups, the last Kalandar dancing bear was set free. The effort to end the practice involved helping the bear handlers find jobs and education, which enabled them to reduce their reliance on dancing-bear income. == Cultural references ==

Charles Catton included the bear in his 1788 book Animals Drawn from Nature and Engraved in Aqua-tinta, describing it as an "animal of the bear-kind" and saying it was properly called the "Petre Bear". In Rudyard Kipling's The Jungle Book, Baloo "the sleepy old brown bear" teaches the Law of the Jungle to the wolf cubs of the Seeonee wolf pack, as well as to his most challenging pupil, the "man-cub" Mowgli. Robert Armitage Sterndale, from whom Kipling derived most of his knowledge of Indian fauna, used the Hindustani word bhalu for several bear species, though Daniel Karlin, who edited the Penguin Classics reissue of The Jungle Book in 1989, stated, with the exception of colour, Kipling's descriptions of Baloo are consistent with the sloth bear, as brown bears and Asian black bears do not occur in the Seoni area where the novel takes place. Also, the name "sloth" can be used in the context of sleepiness. Karlin states, however, that Baloo's diet of ".. only roots and nuts and honey" is a trait more common to the Asian black bear than to the sloth bear. Local names: • , bhaluk • , rīn̄ch; also rinchh • , bhālū; • , bhālu • , ślath bhaluk; kālō bhāluk; also bhaluk • , ṛkṣa; also rikspa • , karaḍi; kaddi • , karaṭi; kaddi • , karaṭi • , elugubaṇṭi; also elugu • , asval; also aswal • Gond: yerid, yedjal and asol • Kol: bana • Oraon: bir mendi • , valasā; also usa • , bhālu • , richh == References ==