'', on an
American quarter dollar Size The now
extinct Titanoboa cerrejonensis was in length. By comparison, the largest
extant snakes are the
reticulated python, measuring about long, At the other end of the scale, the smallest extant snake is
Leptotyphlops carlae, with a length of about . Most snakes are fairly small animals, approximately in length.
Perception The sensory systems of snakes, particularly those of the Crotalidae family, commonly known as pit vipers, are among the most specialized in the animal kingdom. Pit vipers, which include rattlesnakes and related species, possess all the sensory organs found in other snakes, as well as additional adaptations. These include specialized
infrared-sensitive receptors, known as pits, located on either side of the head between the nostrils and eyes. These pits, which resemble an additional pair of nostrils, are highly developed and allow pit vipers to detect minute temperature changes. Each pit consists of two cavities: a larger outer cavity positioned just behind and below the nostril, and a smaller inner cavity. These cavities are connected internally by a membrane containing nerves highly sensitive to thermal variations. The forward-facing pits create a combined field of detection, enabling pit vipers to distinguish objects from their surroundings and accurately judge distances. The sensitivity of these pits allows them to detect temperature differences as small as one-third of a degree Fahrenheit. Other infrared-sensitive snakes, such as those in the Boidae family, possess multiple smaller labial pits along the upper lip, just below the nostrils. Snakes rely heavily on their sense of smell to track prey. They collect particles from the air, ground, or water using their
forked tongue, which are then transferred to the
vomeronasal organ (also known as Jacobson's organ) in the mouth for analysis. The forked structure of the tongue provides directional information of smell which helps locate prey or predators. In aquatic species, such as the
anaconda, the tongue functions efficiently underwater. When the tongue is retracted, the forked tips are pressed into the cavities of the Jacobson's organ, enabling a combined taste-smell analysis that provides the snake with detailed information about its environment. '' by
G. A. Boulenger (1890), illustrating the terminology of shields on the head of a snake Until the mid-20th century, it was widely believed that snakes were unable to hear. However, snakes possess two distinct auditory systems. One system, the somatic system, involves the transmission of vibrations through ventral skin receptors to the spine. The other system involves vibrations transmitted through the snake's elongated lung to the brain via cranial nerves. Snakes exhibit high sensitivity to vibrations, allowing them to detect even subtle sounds, such as soft speech, in quiet environments. Arboreal snakes generally have better vision than burrowing species. Some snakes, such as the
Asian vine snake, possess
binocular vision, enabling both eyes to focus on the same point. Most snakes focus by moving the
lens back and forth relative to the
retina. Diurnal snakes typically have round pupils, while many nocturnal species have slit pupils. Most snakes possess three visual pigments, allowing them to perceive two primary colors in daylight. Certain species, such as the
annulated sea snake and members of the genus
Helicops, have regained significant color vision as an adaptation to their aquatic environments. Research suggests that the last common ancestor of all snakes had
UV-sensitive vision. However, many diurnal snakes have evolved lenses that filter out UV light, likely improving
contrast and sharpening their vision.
Skin The skin of a snake is covered in
scales. Contrary to the popular notion of snakes being slimy (because of possible confusion of snakes with
worms),
snakeskin has a smooth, dry texture. Most snakes use specialized belly scales to travel, allowing them to grip surfaces. The body scales may be smooth,
keeled, or granular. The eyelids of a snake are transparent "spectacle" scales, also known as
brille, which remain permanently closed. For a snake, the skin has been modified to its specialized form of locomotion. Between the inner layer and the outer layer lies the dermis, which contains all the pigments and cells that make up the snake's distinguishing pattern and color. The epidermis, or outer layer, is formed of a substance called
keratin, which in mammals is the same basic material that forms nails, claws, and hair. The snake's epidermis of keratin provides it with the armor it needs to protect its internal organs and reduce friction as it passes over rocks. Parts of this keratin armor are rougher than others. The less restricted portion overlaps the front of the scale beneath it. Between them lies a folded back connecting material, also of keratin, also part of the epidermis. This folded back material gives as the snake undulates or eats things bigger than the circumference of its body. The shedding of scales is called
ecdysis (or in normal usage,
molting or
sloughing). Snakes shed the complete outer layer of skin in one piece. Snake scales are not discrete, but extensions of the
epidermis—hence they are not shed separately but as a complete outer layer during each molt, akin to a sock being turned inside out. Snakes have a wide diversity of skin coloration patterns which are often related to behavior, such as the tendency to have to flee from predators. Snakes that are at a high risk of predation tend to be plain, or have longitudinal stripes, providing few reference points to predators, thus allowing the snake to escape without being noticed. Plain snakes usually adopt active hunting strategies, as their pattern allows them to send little information to prey about motion. Blotched snakes usually use ambush-based strategies, likely because it helps them blend into an environment with irregularly shaped objects, like sticks or rocks. Spotted patterning can similarly help snakes to blend into their environment. The shape and number of scales on the head, back, and belly are often characteristic and used for taxonomic purposes. Scales are named mainly according to their positions on the body. In "advanced" (
Caenophidian) snakes, the broad belly scales and rows of
dorsal scales correspond to the
vertebrae, allowing these to be counted without the need for
dissection.
Molting shedding its skin
Molting (or "ecdysis") serves a number of purposes - it allows old, worn skin to be replaced and can be synced to mating cycles, as with other animals. Molting occurs periodically throughout the life of a snake. Before each molt, the snake regulates its diet and seeks defensible shelter. Just before shedding, the skin becomes grey and the snake's eyes turn silvery. The inner surface of the old skin liquefies, causing it to separate from the new skin beneath it. After a few days, the eyes clear and the snake reaches out of its old skin, which splits. The snake rubs its body against rough surfaces to aid in the shedding of its old skin. In many cases, the castaway skin peels backward over the body from head to tail in one piece, like taking the dust jacket off a book, revealing a new, larger, brighter layer of skin which has formed underneath. Renewal of the skin by molting supposedly increases the mass of some animals such as insects, but in the case of snakes this has been disputed. Shedding skin can release pheromones and revitalize color and patterns of the skin to increase attraction of mates. Snakes may shed four or five times a year, depending on the weather conditions, food supply, age of the snake, and other factors. One can attempt to identify the sex of a snake when the species is not distinctly
sexually dimorphic by counting scales. The
cloaca is probed and measured against the
subcaudal scales. Counting scales determines whether a snake is a male or female, as the
hemipenis of a male being probed is usually longer. In a snake's skull the brain is well protected. As brain tissues could be damaged through the palate, this protection is especially valuable. The solid and complete
neurocranium of snakes is closed at the front. here, for example), consisting almost entirely of an extended ribcage. The skeleton of most snakes consists solely of the skull,
hyoid, vertebral column, and ribs, though
henophidian snakes retain vestiges of the pelvis and rear limbs. The hyoid is a small bone located posterior and ventral to the skull, in the 'neck' region, which serves as an attachment for the muscles of the snake's tongue, as it does in all other
tetrapods. The vertebral column consists of between 200 and 400 vertebrae, or sometimes more. The body vertebrae each have two ribs articulating with them. The tail vertebrae are comparatively few in number (often less than 20% of the total) and lack ribs. The vertebrae have projections that allow for strong muscle attachment, enabling locomotion without limbs. Caudal
autotomy (self-amputation of the tail), a feature found in some lizards, is absent in most snakes. In the rare cases where it does exist in snakes, caudal autotomy is intervertebral (meaning the separation of adjacent vertebrae), unlike that in lizards, which is intravertebral, i.e. the break happens along a predefined fracture plane present on a vertebra. In some snakes, most notably boas and pythons, there are vestiges of the hindlimbs in the form of a pair of
pelvic spurs. These small, claw-like protrusions on each side of the cloaca are the external portion of the vestigial hindlimb skeleton, which includes the remains of an ilium and femur. Snakes are
polyphyodonts with teeth that are continuously replaced.
Internal organs Snakes and other non-
archosaur (
crocodilians,
dinosaurs +
birds and allies) reptiles have a three-chambered heart that controls the
circulatory system via the left and right atrium, and one ventricle. Internally, the ventricle is divided into three interconnected cavities: the cavum arteriosum, the cavum pulmonale, and the cavum venosum. The cavum venosum receives deoxygenated
blood from the right atrium and the cavum arteriosum receives oxygenated blood from the left atrium. Located beneath the cavum venosum is the cavum pulmonale, which pumps blood to the pulmonary trunk. The snake's heart is encased in a sac, called the
pericardium, located at the
bifurcation of the
bronchi. The heart is able to move around, owing to the lack of a diaphragm; this adjustment protects the heart from potential damage when large ingested prey is passed through the
esophagus. The
spleen is attached to the
gall bladder and
pancreas and filters the blood. The
thymus, located in fatty tissue above the heart, is responsible for the generation of immune cells in the blood. The cardiovascular system of snakes is unique for the presence of a renal portal system in which the blood from the snake's tail passes through the kidneys before returning to the heart. In the majority of species, only one lung is functional. This lung contains a vascularized anterior portion and a posterior portion that does not function in gas exchange. The fangs of 'advanced' venomous snakes like viperids and elapids are hollow, allowing venom to be injected more effectively, and the fangs of
rear-fanged snakes such as the boomslang simply have a groove on the posterior edge to channel venom into the wound. Snake venoms are often prey-specific, and their role in self-defense is secondary. According to this theory, most snakes that are labelled "nonvenomous" would be considered harmless because they either lack a venom delivery method or are incapable of delivering enough to endanger a human. The theory postulates that snakes may have evolved from a common lizard ancestor that was venomous, and also that venomous lizards like the
gila monster,
beaded lizard,
monitor lizards, and the now-extinct
mosasaurs, may have derived from this same common ancestor. They share this "
venom clade" with various other
saurian species. Venomous snakes are classified in two taxonomic families: •
Elapids –
cobras including
king cobras,
kraits,
mambas,
Australian copperheads,
sea snakes, and
coral snakes. •
Viperids – vipers,
rattlesnakes,
copperheads/
cottonmouths, and
bushmasters. There is a third family containing the
opistoglyphous (rear-fanged) snakes (as well as the majority of other snake species): •
Colubrids –
boomslangs,
tree snakes,
vine snakes,
cat snakes, although not all colubrids are venomous. The hemipenes are often grooved, hooked, or spined—designed to grip the walls of the female's
cloaca. Most species of snakes lay
eggs which they abandon shortly after laying. However, a few species (such as the king cobra) construct nests and stay in the vicinity of the hatchlings after incubation. Several species of snake, such as the
boa constrictor and green anaconda, are fully
viviparous, nourishing their young through a
placenta as well as a
yolk sac; this is highly unusual among reptiles, and normally found in
requiem sharks or
placental mammals. Ritual combat between males for the females they want to
mate with includes topping, a behavior exhibited by most viperids in which one male will twist around the vertically elevated fore body of its opponent and force it downward. It is common for neck-biting to occur while the snakes are entwined.
Facultative parthenogenesis Parthenogenesis is a natural form of reproduction in which growth and development of embryos occur without fertilization.
Agkistrodon contortrix (copperhead) and
Agkistrodon piscivorus (cottonmouth) can reproduce by
facultative parthenogenesis, meaning that they are capable of switching from a
sexual mode of reproduction to an
asexual mode. The most likely type of parthenogenesis to occur is
automixis with terminal fusion, a process in which two terminal products from the same
meiosis fuse to form a diploid
zygote. This process leads to genome-wide
homozygosity, expression of deleterious recessive
alleles, and often to developmental abnormalities. Both captive-born and wild-born copperheads and cottonmouths appear to be capable of this form of parthenogenesis.
Embryonic development embryo 12 day post
fertilization side by side with
corn snake embryo 2 days post ovo-positioning Cell division and proliferation continues until an early snake embryo develops and the typical body shape of a snake can be observed. This increase in vertebrae is due to an increase in
somites during embryogenesis, leading to an increased number of vertebrae which develop. There is ample literature focusing on the limb development/lack of development in snake embryos and the gene expression associated with the different stages. In
basal snakes, such as the python, embryos in early development exhibit a hind
limb bud that develops with some cartilage and a cartilaginous pelvic element, however this degenerates before hatching. This presence of vestigial development suggests that some snakes are still undergoing hind limb reduction before they are eliminated. There is no evidence in basal snakes of forelimb rudiments and no examples of snake forelimb bud initiation in embryo, so little is known regarding the loss of this trait. == Behavior and life history ==