The series was used during the
Korean War by Chinese and
North Korean forces, and was later considered to be the most dangerous opposition to U.S.
helicopters in
Vietnam. Later it was used by
Morocco and the
Polisario Front in the
Western Sahara War. It was also used by
Iraqi forces during
Operation Desert Storm and again in Operation Iraqi Freedom. In 1974 the Cyprus National Guard artillery batteries used their ZPU-2s against the Turkish air force. In the Russian military, it was replaced by the newer and more powerful
ZU-23 23 mm twin automatic anti-aircraft gun. During the
Lebanese Civil War, the
Lebanese militias mounted the ZPU-2 and ZPU-4 on various vehicles, such as
M113 armored personnel carriers, to create self-propelled support vehicles. with ZU-2. The ZPU has seen widespread use by both sides in the
Libyan Civil War,
Syrian Civil War, and
Yemeni Civil War, being often mounted on
technical pickup trucks. The weapon is credited for bringing down several
Syrian Air Force helicopters. In North Korea, ZPU systems have been modified to be able to be directed by a MR-104 "Drum Tilt", where the guns are shown to be fired without personnel manning them. Several ZPU-4 weapons are suspected to have been used in public executions in the country in October 2014. During the
Donbas War, ZPUs were observed in use with Ukrainian and
pro-Russian separatists forces. Czech company Excalibur Army developed the MR-2 Viktor mobile gun air defense system consisting of a ZPU-2 equipped with day/night sights mounted on a pickup truck chassis upgraded with a reinforced suspension and modified body. The Viktor was supplied to Ukraine in late 2022 during the
Russian invasion of Ukraine to combat low-altitude aerial targets, particularly
unmanned aerial vehicles. ==Ammunition==