portrays
Temperance "Bones" Brennan portrays
Seeley Booth portrays Angela Montenegro portrays
Jack Hodgins portrays
Lance Sweets Main cast •
Emily Deschanel as
Temperance "Bones" Brennan: A brilliant forensic anthropologist, she works at the renowned Jeffersonian Institute located in Washington, D.C. Her birth name was Joy Keenan, which her parents changed after they turned away from a life of crime, and she was educated at
Northwestern University. She is an
empiricist and a wealthy author of
crime fiction drawn from her experiences. She is an atheist and a staunch believer in facts and evidence, to the exclusion of feelings; consequently, she comes off as distant and detached. Her lack of social skills provides most of the show's lighthearted humor, primarily through her catchphrase, "I don't know what that means," whenever a
pop culture reference is introduced into the conversation. She is noted to have a high IQ and impressive reasoning skills. Despite being superficially very different, Booth and she share a deep connection and begin a relationship near the end of season six. In season seven, they have a daughter together, whom they name Christine, in honor of Temperance's late mother. They marry in season nine. Their son, Hank, named after Booth's paternal grandfather, is born in season 11. She is also stepmother to Booth's son from a previous relationship, Parker. •
David Boreanaz as
Seeley Booth: FBI Special Agent Booth seeks out Brennan's professional help in his investigations involving human remains that cannot be identified without her skills. His character is often used as an
audience surrogate to provide a layman's translation of the scientific jargon-filled dialogue, especially in the "squints" conversations, or laboratory scenes. He gives Brennan her nickname, "Bones", which she first hates, but comes to accept. He is a skilled investigator and interrogator who often relies on his "gut" and "cop instincts" (qualities foreign to Brennan). He is a decorated veteran of the
United States Army Rangers, where he was a
sniper. Booth has a son, Parker, from a previous relationship, and a younger brother, Jared, an officer in the
U.S. Navy. He has two children with Brennan: a daughter, Christine, and a son, Hank. Raised in the
Roman Catholic church, Booth is a devout believer in God and heaven. He frequently debates spiritual issues with Brennan, who regards all religions as primitive superstitions unsupported by empirical evidence. He plays hockey on a team with intern
Wendell Bray, with whom he is close. He is purportedly a member of the well-known
Booth family, and as such is related to Lincoln's assassin
John Wilkes Booth. •
Michaela Conlin as Angela Montenegro: A
forensic artist at the Jeffersonian Institute and Brennan's best friend, Montenegro is also her team specialist in
forensic facial reconstruction—helping
identify the victims. She can also generate holograms using her three-dimensional graphics program (the "Angelator" and later, the "Angelatron") to simulate various scenarios of a crime to help determine the cause of death. She and Brennan are best friends and rarely argue. She is open, friendly, and caring and repeatedly tries to draw Brennan out of the lab. She is often described as a free spirit. Angela's father is played by
Billy Gibbons, guitarist of
ZZ Top, who guest stars as a fictional version of himself. In season two, Angela begins a relationship with Jack Hodgins. They try to wed, but Angela discovers she is already married to a man she barely remembers; after her divorce from him is finalized, she and Hodgins break up. During this time, Angela dates intern Wendell Bray, but she marries Hodgins in season five and gives birth to their son, Michael Vincent, in season six. Her birth name is Pookie Noodlin, as revealed in the 10th season. •
Eric Millegan as
Dr. Zack Addy (seasons 1–3; guest star, seasons 4–5, 11–12) is introduced in season one as Brennan's graduate student and intern. In season two, he receives his
doctorates in forensic anthropology and applied engineering, and joins the staff of the Jeffersonian as a full-fledged professional. Like Brennan, his inability to pick up on Booth, Hodgins, and Montenegro's pop-culture jokes and references is a running gag and a source of humorously awkward moments in the lab, though he has shown he has other "normal" interests such as being a
basketball fan and a trained singer. A recurring element in the show involves Addy and Hodgins amicably but resolutely competing to be "King of the Lab". In the episode, "The Pain in the Heart", a trophy, presented to him by Saroyan that recognizes Zack as "King of the Lab", is revealed. Zack is removed from his position on Brennan's team in the season three finale, "The Pain in the Heart", when he is revealed to be the apprentice of the serial killer
Gormogon. He resides in a psychiatric facility after pleading insanity to avoid a prison term, as Hodgins told him he "would not do well" in the general prison population. At the end of season 11, Zack escapes the facility after he is suspected of being another serial killer called "The Puppeteer". At the beginning of season 12, Zack is exonerated when his doctor is proven to be the killer, instead. Zack confesses his innocence in the Gormogon murders to Booth and Brennan. A major storyline in season 12 is exonerating Zack. In the penultimate episode, "A Day in the Life", Zack is exonerated of murder thanks to Hodgins, with help from Brennan and Caroline Julian at his hearing. However, the judge rules that he must finish out the last 13 months of his sentence for the separate crime of aiding a known killer. Zack remains institutionalized, albeit with a release date just over a year away. •
T. J. Thyne as
Jack Hodgins is an
entomologist who is also an expert on
spores and
minerals, but whose
hobby is
conspiracy theories. Hodgins regularly identifies as the lab's "bug and slime guy". During an investigation, he primarily deals with
particulates and
trace evidence, and at the crime scene will provide Booth with an approximate time of death. He is best friends with former coworker Zack Addy and often tries to "educate" him on
social norms. They often compete for the title of "King of the Lab", referring to whoever has more useful information in the investigation at hand. His family owns the fictional Cantilever Group, and he is extremely wealthy and is a major sponsor of the Jeffersonian. However, Hodgins tries to keep this concealed from his colleagues. It eventually comes out during an investigation, but no one allows his family's wealth to significantly affect their relationship with him. He later loses all his money due to the scheming of serial killer
Christopher Pelant, but in time, Hodgins regains a considerable portion of it. He sells a hot sauce he develops in collaboration with intern Finn Abernathy, which becomes immensely popular. Later, he develops an unbreakable beaker and a polymer floor mat. He marries Angela Montenegro in season five. She gives birth to their son, Michael Vincent, in season six. In season 11, Hodgins is wounded in a bomb attack and left paralyzed from the waist down. At the end of the series, Hodgins is promoted to temporary head of the Jeffersonian lab while Cam and Arastoo are on leave after their wedding. •
Jonathan Adams as
Daniel Goodman (season 1): An
archaeologist turned administrator, he is the director of the Jeffersonian Institute. He is a loving husband and father to a pair of five-year-old twin girls. His way of working leads Hodgins to think of him as
subjective, long-winded, and lacking the qualities of a pure scientist, although the antagonism between the two develops into a friendly rivalry as the season progresses. He has not made any appearances beyond season one. As of episode 23, "The Titan on the Tracks", he is said to be on
sabbatical, and is not seen for the rest of the series. •
Tamara Taylor as
Camille Saroyan (seasons 2–12) succeeds Dr. Goodman as the head of the forensic division at Jeffersonian Institute and is a
pathologist. She was a coroner in New York City. Cam had a romantic relationship with Booth before she joined the Jeffersonian, and a brief relationship during the show. At first, Brennan and she have an uneasy working relationship, but she grows to be an integral member of the Jeffersonian team and well respected by all of its members, including Brennan. In season four, she adopts 16-year-old Michelle, whose father Cam previously dated and lived with, after Michelle's father is murdered. Later in the series, she becomes romantically involved with intern Arastoo Vaziri. They marry in the series' penultimate episode "The Day in the Life", taking a sabbatical after the final case is concluded. •
John Francis Daley as
Lance Sweets (seasons 3–10) is an FBI
psychologist assigned to Booth and Brennan after Booth arrests her father. He is frequently consulted to provide a profile of the suspects and victims involved and to give a more "humanized" perspective on the case. Due to his age, he is initially treated by Booth and the Jeffersonian team in a condescending manner, but eventually earns their respect and their friendship; another FBI psychologist notes to Brennan and Booth that Sweets has imprinted onto them like a duckling, and he briefly lived with them. Throughout the series, he has an on-again/off-again relationship with intern
Daisy Wick. He eventually is accepted by Brennan, who notably dismisses psychology as a "soft science", and is often sought out for advice by the team on both professional and personal matters. In the season 10 premiere, Daisy and he are revealed to be expecting a baby. Soon after, he is fatally assaulted while serving a warrant. He dies with Booth and Brennan by his side. •
John Boyd as
James Aubrey (seasons 10–12) is a junior FBI agent working under Booth after Sweets's death. Originally, Aubrey tries to win Booth's approval and trust, as Booth is reluctant to work with him due to Aubrey's inexperience and his grief for Sweets. However, he is eventually accepted by Booth and the rest of the team. Aubrey gets along with everyone, and his love of food is often the subject of jokes. When he was a child, his father was an investment broker who cheated his clients and fled the country, abandoning Aubrey and his mother and leaving them with nothing. In season 11, Aubrey's father is found to be back in the country. In season 12, Aubrey is finally approached by his father, who asks him for money. He reveals he has another wife and young son back in the country where he had fled, to persuade Aubrey to give him financial aid. However, Aubrey realizes it is all a lie and eventually turns his father in. Near the end of season 12, Aubrey is offered a promotion to supervisory special agent in Los Angeles. Though Aubrey originally took the Los Angeles job, he instead decides to take a similar position in Washington, D.C., in the series finale.
Recurring cast •
Loren Dean as Russ Brennan (seasons 1–3): Russ Brennan, or Kyle Keenan, is the older brother of Temperance Brennan - Joy Keenan, and the son of Max Keenan. Russ has a troubled past and has spent some time in prison. He is involved with a woman named Amy Hollister and loves her two daughters, Emma and Hayley. After Temperance and he were abandoned by their parents, he also abandoned his sister, which led to long-held animosity from her. He first appears in the season one finale, where he patches things up with his sister and also reunites with his father in season two. His last appearance is in season three, although he is mentioned in later seasons. •
Billy Gibbons as Angela Montenegro's father (seasons 1–10): The character is a fictionalized version of Gibbons himself–a bearded guitarist in a famous rock band that plays sold-out stadium shows–but despite all the knowing winks to him being Gibbons, the character is never named, only referred to in dialogue as "Angela's father", "Dad," "my father". •
Patricia Belcher as
Caroline Julian (seasons 1–12): Caroline is the acerbic, witty, and often meddlesome
assistant U.S. attorney (AUSA) from the
U.S. Department of Justice. She is a friend of Booth's and is always willing, if not eager, to help his partner and him in their cases. She gives Booth a hard time after he accidentally wrecks her prized
AMC Gremlin in the course of a hot pursuit. •
Ryan O'Neal as Max Keenan (seasons 2–12): Max Keenan, or Matthew Brennan, is the father of Russ and Temperance Brennan. His wife and he were nonviolent bank robbers who became involved with gangs in the 1970s. They changed their identities to lead honest lives with their children. Booth arrests Max on murder charges in season two, and he is tried in season three. The jury finds him not guilty on all counts and he is released, finally able to reconnect openly with his children. He is a frequent presence in Brennan and Booth's lives, particularly after the birth of his granddaughter Christine. During season 12, he dies while protecting his grandchildren. •
Eugene Byrd as Clark Edison (seasons 3–12) is the most serious and professional of the Jeffersonian interns, as well as the most organized; he initially dislikes discussing his personal life and his coworkers' personal lives while at work. While Brennan is on the run after being suspected of murder, Cam hires Clark as the new forensic anthropologist. After Brennan's return, Cam keeps him on to focus on archaeological work while Brennan works on criminal investigations. •
Brendan Fehr as Jared Booth (seasons 4–5) is Agent Seeley Booth's younger brother, who is notorious for getting into trouble. However, in a continuing "Gravedigger" episode, Jared uses his military contacts to help save his brother's life, at the cost of his Navy career. In seasons four and five, his appearances are troublesome to his older brother, especially when he announces his engagement to a former escort, whom he later divorces. In season 11, the trend continues, but this time with a much more dire outcome, as Jared is the newest victim for the Jeffersonian. Detached forensic work on Brennan's part is needed to discover that his remains are not those of Seeley Booth. •
Ryan Cartwright as Vincent Nigel-Murray (seasons 4–6) is an Englishman and a graduate of the
University of Leeds. He has a habit of reciting trivia while working. In season six, he is fatally shot in the chest by sniper Jacob Broadsky; his death ends up being the catalyst in Booth and Brennan's relationship. Hodgins and Angela give their son the second middle name Vincent. •
Michael Grant Terry as Wendell Bray (seasons 4–12) is Dr. Brennan's top scholarship intern. He dates Angela briefly during season five and is close friends with Booth, who regards Wendell as the most relatable of the interns. In season nine, he is diagnosed with
Ewing's sarcoma by Brennan after breaking his arm during a hockey game. After seeking treatment, he appears to be in full remission and continues to work at the lab. In late season 12, after Wendell has trouble writing his thesis, Brennan helps him realize that his passion is not forensic anthropology after all. By the end of the series, Wendell intends to leave the Jeffersonian to find his true passion. •
Carla Gallo as Daisy Wick (seasons 4–12): Overeager and somewhat manipulative, she is often annoying to other workers at the Jeffersonian, though she eventually develops a close friendship with Hodgins. She has an
on-again, off-again relationship with Lance Sweets. Sweets also helps Daisy and the others in the lab get along with each other. They later have a son named Seeley Lance Wick-Sweets, born some months after his father's death. •
Joel David Moore as Colin Fisher (seasons 4–12) has depression and has a very dark sense of humor. He sometimes spends time at mental-health facilities for treatment, so is the intern least often shown working in the lab. He is adept at estimating bone remodeling. •
Pej Vahdat as Arastoo Vaziri (seasons 4–12) starts by having a fake Iranian accent, so he does not have to constantly explain his Islamic beliefs, but later admits it is fake. He is a sensitive and devout man, and begins dating Cam in season eight; they become engaged in season 11 and get married in season 12. •
Luke Kleintank as Finn Abernathy (seasons 7–9) is the youngest intern to date, and has a messy background that includes a stay at a juvenile detention center. He is from
North Carolina and has a thick
Southern accent, which leads Hodgins to call him "
Opie". Finn, in turn, calls Hodgins, who is from a very wealthy family, "
Thurston". Hodgins and he go into business together to sell a hot sauce that Abernathy's grandmother made before she died, after Hodgins uses traces from Abernathy's last bottle of the sauce to reverse-engineer the recipe. Cam's daughter Michelle and he fall in love and begin dating, initially behind her mother's back. She later breaks up with him after moving away to college, realizing that the distance and new developments in their lives have caused them to drift apart. •
Brian Klugman as Oliver Wells (seasons 8–11) is a lab assistant who arrogantly boasts of his 160 IQ. He has multiple doctorates in various subjects and is the only intern who is a physician. He struggles to get along with all of the lab members. Brennan and he butt heads, especially after he tells her that he aims to surpass her reputation. A self declared
polymath, he has a wide, if indolent, range of interests, from physics to psychology. •
Laura Spencer as Jessica Warren (seasons 9–12) grew up in an educational cooperative. She has five older brothers and she graduated from Michigan State University top of her class at the age of 19. She desires creativity in science, which contrasts with Brennan's insistence upon no speculation. She believes in trusting her gut. She is somewhat outspoken. In season 9, she first appears in the episode "The Drama in the Queen". At the end of the episode, she has a one-night stand with Sweets. In season 11, she engages in a relationship with FBI agent James Aubrey, which ends in season 12. •
Ignacio Serricchio as Rodolfo Fuentes (seasons 9–12) is a Cuban forensic anthropologist seeking asylum in the U.S. While he was already a qualified forensic anthropologist in Cuba, he is working towards the same qualifications in America. He is also a shameless flirt and has openly flirted with Angela and Brennan, both of whom are married. Though Angela was flattered by his attentions, both women turned down his advances. Despite his flirting, Hodgins takes him under his wing and becomes a good friend to him. He carries his father's prayer beads with him despite not believing in God because he believes that one needs to experience a little bit of everything to make good life decisions. While starting as an intern, Fuentes eventually gets his doctorate and joins the Jeffersonian staff in season 12. •
Sara Rue as Karen Delfs (seasons 11–12) is a flaky behavioral analyst assigned to help Booth and Bones on a case. She asked Agent Aubrey out, but he was seeing Jessica Warren at the time. She transferred to Kansas after a few episodes, but she returned in the first episode of season 12 after being sexually harassed at that job by her married boss. She was accused of being the one out to kill Bones (she had the requisite psychological skills), but was cleared when they discovered the real killer. In the second-to-last episode, Aubrey got falling-down drunk at Cam and Arastoo's wedding reception. Karen had Jessica bring him to her apartment to recover, since it was closer to the reception hall. In the final episode, she learned that Aubrey and Jessica broke up. She bought him some fried chicken and they went off to eat comfort food and commiserate. The implication was that they were on the verge of starting a relationship. ==Production==