Beaver

The English word beaver comes from the Old English word or and is connected to the German word and the Dutch word . The ultimate origin of the word is an Indo-European root for . Cognates of beaver are the source for several European placenames, including those of Beverley, Bièvres, Biberbach, Biebrich, Bibra, Bibern, Bibrka, Bobr, Bober, Bóbrka, Bjurholm, Bjurälven, and Bjurum. The genus name Castor has its origin in the Greek word and translates as . == Taxonomy ==

Carl Linnaeus coined the genus name Castor as well as the specific (species) epithet fiber for the Eurasian species. German zoologist Heinrich Kuhl coined C. canadensis in 1820. Many scientists considered both names synonymous for one same species Twenty-five subspecies have been classified for C. canadensis, and nine have been classified for C. fiber. In the early Miocene (about 24 mya), castorids evolved a semiaquatic lifestyle. Members of the subfamily Castoroidinae are considered to be a sister group to modern beavers, and included giants like Castoroides of North America and Trogontherium of Eurasia. Fossils of one genus in Castoroidinae, Dipoides, have been found near piles of chewed wood, The genus Castor likely originated in Eurasia. Mitochondrial DNA studies place the common ancestor of the two living species at around 8 mya. The ancestors of the North American beaver would have crossed the Bering Land Bridge around 7.5 mya. Castor may have competed with members of Castoroidinae, which led to niche differentiation. The fossil species C. praefiber was likely an ancestor of the Eurasian beaver. C. californicus from the Early Pleistocene of North America was similar to but larger than the extant North American beaver.' ==Characteristics==

Beavers are the second-largest living rodents, after capybaras. They have a head–body length of , with a tail, a shoulder height of , and generally weigh , but can be as heavy as . Males and females are almost identical externally. Their bodies are streamlined like marine mammals and their robust build allows them to pull heavy loads. The lower incisors have roots that are almost as long as the entire lower jaw. Beavers have one premolar and three molars on all four sides of the jaws, adding up to 20 teeth. The molars have meandering ridges for grinding woody material. The eyes, ears and nostrils are arranged so that they can remain above water while the rest of the body is submerged. The nostrils and ears have valves that close underwater, while nictitating membranes cover the eyes. To protect the larynx and trachea from water flow, the epiglottis is contained within the nasal cavity instead of the throat. In addition, the back of the tongue can rise and create a waterproof seal. A beaver's lips can close behind the incisors, preventing water from entering their mouths as they cut and bite onto things while submerged. On the surface, the hind limbs thrust one after the other; while underwater, they move at the same time. Beavers are awkward on land but can move quickly when they feel threatened. They can carry objects while walking on their hind legs. When diving, their heart rate decreases to 60 beats per minute, half its normal pace, and blood flow is directed more towards the brain. A beaver's body also has a high tolerance for carbon dioxide. When surfacing, the animal can replace 75 percent of the air in its lungs in one breath, compared to 15 percent for a human. ==Distribution and status==

. The IUCN Red List of mammals lists both beaver species as least concern. The North American beaver is widespread throughout most of the United States and Canada and can be found in northern Mexico. The species was introduced to Finland in 1937 (and then spread to northwestern Russia) and to Tierra del Fuego, Patagonia, in 1946. Historically, the North American beaver was trapped and nearly extirpated because its fur was highly sought after. Protections have allowed the beaver population on the continent to rebound to an estimated 6–12million by the late 20th century; still far lower than the originally estimated 60–400million North American beavers before the fur trade. The introduced population in Tierra del Fuego is estimated at 35,000–50,000 individuals . , the total Eurasian beaver population in Europe was estimated at over one million. Small native populations are also present in Mongolia and northwestern China; their numbers were estimated at 150 and 700, respectively, . ==Ecology==

Beavers live in freshwater ecosystems such as rivers, streams, lakes and ponds. Water is the most important component of beaver habitat; they swim and dive in it, and it provides them refuge from land predators. It also restricts access to their homes and allows them to move building objects more easily. Beavers prefer slower moving streams, typically with a gradient (steepness) of one percent, though they have been recorded using streams with gradients as high as 15 percent. Beavers are found in wider streams more often than in narrower ones. They also prefer areas with no regular flooding and may abandon a location for years after a significant flood. Beavers typically select flat landscapes with diverse vegetation close to the water. North American beavers prefer trees being or less from the water, but will roam several hundred meters to find more. Beavers have also been recorded in mountainous areas. Dispersing beavers will use certain habitats temporarily before finding their ideal home. These include small streams, temporary swamps, ditches, and backyards. These sites lack important resources, so the animals do not stay there permanently. Beavers have increasingly settled at or near human-made environments, including agricultural areas, suburbs, golf courses, and shopping malls. Food and feeding Beavers have an herbivorous and a generalist diet. During the spring and summer, they mainly feed on herbaceous plant material such as leaves, roots, herbs, ferns, grasses, sedges, water lilies, water shields, rushes, and cattails. During the fall and winter, they eat more bark and cambium of woody plants; tree and shrub species consumed include aspen, birch, oak, dogwood, willow and alder. while others suggest beavers principally forage based on stem size. Beavers may cache their food for the winter, piling wood in the deepest part of their pond where it cannot be reached by other browsers. This cache is known as a "raft"; when the top becomes frozen, it creates a "cap". When chewing down a tree, beavers switch between biting with the left and right side of the mouth. Tree branches are then cut and carried to their destination with the powerful jaw and neck muscles. Other building materials, like mud and rocks, are held by the forelimbs and tucked between the chin and chest. Beavers start building dams when they hear running water, and the sound of a leak in a dam triggers them to repair it. To build a dam, beavers stack up relatively long and thick logs between the banks. Heavy rocks keep them stable, and grass is packed between them. Beavers continue to pile on more material until the dam slopes in a direction facing upstream. Dams can range in height from to and can stretch from to several hundred meters long. Beaver dams are more effective in trapping and slowly leaking water than man-made concrete dams. Lake-dwelling beavers do not need to build dams. Beavers make two types of lodges: bank lodges and open-water lodges. Bank lodges are burrows dug along the shore and covered in sticks while the more complex freestanding, open-water lodges are built over a platform of piled-up sticks. The lodge is mostly sealed with mud, except for a hole at the top which acts as an air vent. Both types are accessed by underwater entrances. As they drag wood across the land, beavers leave behind trails or "slides", which they reuse when moving new material. In one study, beavers were associated with large increases in open-water areas. When beavers returned to an area, 160% more open water was available during droughts than in previous years, when they were absent. Beaver dams also lead to higher water tables in mineral soil environments and in wetlands such as peatlands. In peatlands particularly, their dams stabilize the constantly changing water levels, leading to greater carbon storage. Beaver ponds, and the wetlands that succeed them, remove sediments and pollutants from waterways, and can stop the loss of important soils. These ponds can increase the productivity of freshwater ecosystems by accumulating nitrogen in sediments. Beavers may contribute to climate change. In Arctic areas, the floods they create can cause permafrost to thaw, releasing methane into the atmosphere. As wetlands are formed and riparian habitats are enlarged, aquatic plants colonize the newly available watery habitat. Another study in semiarid eastern Oregon found that the width of riparian vegetation on stream banks increased several-fold as beaver dams watered previously dry terraces adjacent to the stream. Riparian ecosystems in arid areas appear to sustain more plant life when beaver dams are present. Beaver ponds act as a refuge for riverbank plants during wildfires, and provide them with enough moisture to resist such fires. Introduced beavers at Tierra del Fuego have been responsible for destroying the indigenous forest. Unlike trees in North America, many trees in South America cannot grow back after being cut down. (Oncorhynchus nerka) jumping a beaver dam. Beaver activity impacts communities of aquatic invertebrates. Damming typically leads to an increase of slow or motionless water species, like dragonflies, oligochaetes, snails, and mussels. This is to the detriment of rapid water species like black flies, stoneflies, and net-spinning caddisflies. Beaver floodings create more dead trees, providing more habitat for terrestrial invertebrates like Drosophila flies and bark beetles, which live and breed in dead wood. The presence of beavers can increase wild salmon and trout populations, and the average size of these fishes. These species use beaver habitats for spawning, overwintering, feeding, and as havens from changes in water flow. The positive effects of beaver dams on fish appear to outweigh the negative effects, such as blocking of migration. Beaver ponds have been shown to be beneficial to frog populations by protecting areas for larvae to mature in warm water. The stable waters of beaver ponds also provide ideal habitat for freshwater turtles. Beavers help waterfowl by creating increased areas of water. The widening of the riparian zone associated with beaver dams has been shown to increase the abundance and diversity of birds favoring the water's edge, an impact that may be especially important in semi-arid climates. Fish-eating birds use beaver ponds for foraging, and in some areas, certain species appear more frequently at sites where beavers were active than at sites with no beaver activity. In a study of Wyoming streams and rivers, watercourses with beavers had 75 times as many ducks as those without. As trees are drowned by rising beaver impoundments, they become an ideal habitat for woodpeckers, which carve cavities that may be later used by other bird species. Large herbivores, such as some deer species, benefit from beaver activity as they can access vegetation from fallen trees and ponds. ==Behavior==

Beavers are mainly nocturnal and crepuscular, and spend the daytime in their shelters. In northern latitudes, beaver activity is decoupled from the 24-hour cycle during the winter, and may last as long as 29 hours. They do not hibernate during winter, and spend much of their time in their lodges. Territories and spacing near its dam. Beavers typically disperse from their parental colonies during the spring or when the winter snow melts. They often travel less than , but long-distance dispersals are not uncommon when previous colonizers have already exploited local resources. Beavers are able to travel greater distances when free-flowing water is available. Individuals may meet their mates during the dispersal stage, and the pair travel together. It may take them weeks or months to reach their final destination; longer distances may require several years. Beavers establish and defend territories along the banks of their ponds, which may be in length. Beavers mark their territories by constructing scent mounds made of mud and vegetation, scented with castoreum. Those with many territorial neighbors create more scent mounds. Scent marking increases in spring, during the dispersal of yearlings, to deter interlopers. Beavers are generally intolerant of intruders and fights may result in deep bites to the sides, rump, and tail. Beavers are also more tolerant of individuals that are their kin. They recognize them by using their keen sense of smell to detect differences in the composition of anal gland secretions. Anal gland secretion profiles are more similar among relatives than unrelated individuals. Communication Beavers within a family greet each other with whines. Kits will attract the attention of adults with mews, squeaks, and cries. Defensive beavers produce a hissing growl and gnash their teeth. Tail slaps, which involve an animal hitting the water surface with its tail, serve as alarm signals warning other beavers of a potential threat. An adult's tail slap is more successful in alerting others, who will escape into the lodge or deeper water. Juveniles have not yet learned the proper use of a tail slap, and hence are normally ignored. Eurasian beavers have been recorded using a territorial "stick display", which involves individuals holding up a stick and bouncing in shallow water. ==Interactions with humans==

feeding his beaver.|alt=Black and white photo of a man feeding a beaver Beavers sometimes come into conflict with humans over land use; individual beavers may be labeled as "nuisance beavers". Beavers can damage crops, timber stocks, roads, ditches, gardens, and pastures via gnawing, eating, digging, and flooding. Some of these attacks have been fatal, including at least one human death. Beavers can spread giardiasis ('beaver fever') by infecting surface waters, though outbreaks are more commonly caused by human activity. Flow devices, like beaver pipes, are used to manage beaver flooding, while fencing and hardware cloth protect trees and shrubs from beaver damage. If necessary, hand tools, heavy equipment, or explosives are used to remove dams. Hunting, trapping, and relocation may be permitted as forms of population control and for removal of individuals. The Martinez beavers became famous in the mid-2000s for their role in improving the ecosystem of Alhambra Creek in Martinez, California. Zoos have displayed beavers since at least the 19th century, though not commonly. In captivity, beavers have been used for entertainment, fur harvesting, and for reintroduction into the wild. Captive beavers require access to water, substrate for digging, and artificial shelters. Archibald Stansfeld "Grey Owl" Belaney pioneered beaver conservation in the early 20th century. Belaney wrote several books, and was first to professionally film beavers in their environment. In 1931, he moved to a log cabin in Prince Albert National Park, where he was the "caretaker of park animals" and raised a beaver pair and their four offspring. Dorothy Burney Richards, inspired by Belaney, lived with beavers in her home at Beaversprite. Commercial use Beavers have been hunted, trapped, and exploited for their fur, meat, and castoreum. Since the animals typically stayed in one place, trappers could easily find them and could kill entire families in a lodge. Many pre-modern people mistakenly thought that castoreum was produced by the testicles or that the castor sacs of the beaver were its testicles, and females were hermaphrodites. Aesop's Fables describes beavers chewing off their testicles to preserve themselves from hunters, which is impossible because a beaver's testicles are internal. This myth persisted for centuries, and was corrected by French physician Guillaume Rondelet in the 1500s. Beavers have historically been hunted and captured using deadfalls, snares, nets, bows and arrows, spears, clubs, firearms, and leg-hold traps. Castoreum was used to lure the animals. they preferred its meat more than other red meats because of its higher calorie and fat content, and the animals remained plump in winter when they were most hunted. The bones were used to make tools. The fur trade peaked between 1860 and 1870, when over 150,000 beaver pelts were purchased annually by the Hudson's Bay Company and fur companies in the United States. The contemporary global fur trade is not as profitable due to conservation, anti-fur and animal rights campaigns. ==See also==