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Eusideroxylon

Eusideroxylon is a genus of evergreen trees of the family Lauraceae. The genus is monotypic, and includes one accepted species, Eusideroxylon zwageri. The species is commonly known as Bornean ironwood, or by the Malay names belian and ulin. It is predominantly found in Borneo and Sumatra, where it grows in lowland rain forests, but is also thought to inhabit the Philippines.

Nomenclature
The name Eusideroxylon is Latinised Greek, derived from Greek sideros meaning iron, xylon meaning wood, though some sources retain it in Eusideroxylon. Embryological studies suggest Eusideroxylon clades together with the Cryptocaryeae. ==Description==
Description
Eusideroxylon zwageri is a canopy tree species with erect or spreading branches and extremely durable and decay-resistant wood. They are slow growing ( per year) tall evergreen trees with a straight bole (usually host to Cassytha, a parasitic vine with leaves reduced to scales, up to half of the tree's height). It is slightly fluted at the base, up to in diameter. The trunk has many small, rounded "buttresses" that give the base an elephant-foot like appearance. Common commercially exploitable trees attain a height of around with trunk diameters of exploitable trees up to . Protected trees are towering giants of the forest attaining a height of up to and a diameter of 220 cm – though height is routinely reduced by lightning strikes. An Ulin tree discovered in 1993 in Kutai National Park, is one of the largest plants in Indonesia. It is an estimated 1,000 years old, and has increased its diameter from in the 20 years since its discovery. Its height was however reduced from about to only after a lightning strike. Odoardo Beccari reported a specimen with a girth of 10 metres. The leaves are dark green, simple, leathery, elliptical to ovate, long and wide, and are alternate, rarely whorled or opposite, without stipules and petiolate. The leaf blade is entire (unlobed or lobed in Sassafras) and occasionally with domatia (crevices or hollows serving as lodging for mites) in axils of main lateral veins (present in Cinnamomum). Young leaves are reddish brown to yellowish red. They have a generous layer of wax, making them glossy in appearance, and are narrow, pointed oval in shape with an apical mucro, or 'drip tip', which enables the leaves to shed excess water in a humid environment. The flowers are pale yellow to yellow. The flower is hermaphrodite, actinomorphic, with 6 tepals, distributed in two whorls that overlap. There are six staminodes, three stamens, and a simple pistil that consists of one carpel. Pollination is done by bees and other insects. The fruits are drupes, varying in size and shape from oblong to ovate or sub-cylindrical to asymmetric elongated or rounded. They are long, in diameter, and weigh . Based on morphology, four varieties (var.) are recognized, being exilis, ovoidus, grandis, and zwageri. Generation length is thought to be around 100 years, Genetic variability is relatively high in Indonesia, and populations that are geographically close may be more distant genetically. A number of populations from Kalimantan exhibit evidence of inbreeding. ==Ecology==
Ecology
s Eusideroxylon zwageri seedlings require some shade, while older trees need plenty of light. It can be found in valleys and on hillsides and even on low ridges when soil moisture is sufficient, throughout elevations between sea level and . It may be found in secondary forests and a variety of soil types. hornbills, and raptors. Megafauna such as the Bornean rhino and various extinct species may have dispersed them as well. The seeds may drift out to sea, and have been found on offshore islands such as Maratua. ==Relation to humans==
Relation to humans
The species is considered unsuitable for large-scale plantations due to slow growth and inadequate seed and seedling supply. Manual selection of trees in natural forests is common. The standing timber volume of trees with a diameter of over may be as much as . Properties The wood is dense, and texture is moderately fine to fine and even. Also attractive to users is the resistance to insects, bacteria, fungi and marine borers. The wood has anti-bacterial properties (for local medicinal use) The heartwood when cut is coloured light brown to almost bright yellow. During the aging process the heartwood darkens to deep reddish brown, very dark brown or almost black. The sapwood is bright yellow when cut, and darkens slightly. The wood texture is fine and even, with a straight grain or only slightly interlocked. The timber retains a pleasant lemon odour. This odour, along with the woods' natural high lustre, make it prized by cabinet-makers and fine furniture craftsmen. Vessels are diffuse-porous, medium-sized and generally evenly distributed, arranged in short radial rows (2–3 vessels). Moderate abundance of aliform paratracheal parenchyma. Growth rings boundaries are indistinct or absent. Tyloses are often present. the wood is highly prized for many outdoor uses, especially as decking. Additionally, the wood's high density and easy workability lend it particular desirability in maritime structures, dock construction and ship building, especially Indonesia's pinisi sail-boat. Common local uses include house construction, door construction, water butts and troughs, boat building (such as for pinisi), tools, tool handles, talisman, jewellery, medicinal slivers (for wounds, cuts, abrasions, bites and tooth-ache/infection), bridges, blowpipes and spear shafts, shingles, and flooring. The tree is important to the Dayak and Berwan peoples, and is assigned by them mystical powers such as protection against large animals. Internationally, it is renowned for heavy construction such as a buffer between transportation trailers and heavy steel fabrications (such as boilers, pressure vessels, reactors and many others). It is also frequently found in dry docks as a timber to separate the hull of ships from the steel supporting stands. Other uses include use in boats and ships, industrial flooring, roofing (as shingles), fine indoor and outdoor furniture, coffin wood (esteemed by Chinese due to ability to withstand rot and insect attack) and tool handles (especially those exposed to continual high impact (the wood does not splinter and thus injure hands, eyes or endanger the operator on catastrophic failure) such as shovels, axes, block splitters, sledge hammers, heavy mallets, demolition hammers, mattocks, picks, hoes, and other types of hammer). Some expert cabinet-makers treasure an ulin-headed carpenter's mallet as an excellent intermediate density hammer face between the usual wood and a metal one and is able to quite easily tap or "whack" stubborn highly polished metal fixtures without damage to the face or the fixture. Other sources indicate that ulin wood is often used for marine constructions such as pilings, wharfs, docks, sluices, dams, ships, bridges, but also used for power line poles, masts, roof shingles and house posts and to a minor extent as frame, board, heavy duty flooring, railway sleepers, fencing material, furniture etc. Ironwood extract (essential oils) may be a promising source of antioxidants and termite control substances. Flavonoids have also been extracted from this species. Conservation The government of Indonesia and the state government of Sarawak have formally banned the export of this species. Illegal logging and smuggling continues to be a major problem. Population decline is estimated to be around 30% over the last 300 years. Conservation efforts are underway, with several countries banning imports. ==References==
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