Lily of the valley

Convallaria majalis is a herbaceous perennial plant that often forms extensive colonies by spreading underground stems called rhizomes. New upright shoots are formed at the ends of stolons in summer. The flowering stems have a one-sided raceme of six to twelve pendulous flowers on the upper part of the stem. The flowers have six white tepals (except for the species Convallaria majalis var. "Rosea", which has pink tepals), fused at the base to form a bell shape with reflexed tips that are in diameter, and are sweetly scented; flowers in late spring, typically May to June in Britain; in mild winters in the Northern Hemisphere it can flower as early as March. The fruit is a poisonous small orange-red berry that is approximately in diameter that contains an average of 3.9 large whitish to brownish seeds that dry to a clear translucent round bead wide. The fruit persists for an average of 47.5 days. The fruit averages 85.8% water, and their dry weight includes 14.6% carbohydrates and 1.3% lipids. Plants are self-incompatible, and colonies consisting of a single clone do not set seed. ==Taxonomy==

In the APG III system, the genus is placed in the family Asparagaceae, subfamily Convallarioideae (formerly the family Ruscaceae). It was formerly placed in its own family Convallariaceae, and, like many lilioid monocots, before that in the lily family Liliaceae. In the past, it was widely treated in three varieties, but these are now separated out as distinct species. • Convallaria majalis var. majalis, from Europe and western Asia, with white midribs on the flowers – now known as Convallaria majalis sensu stricto. • Convallaria majalis var. montana – from the southern Appalachian Mountains, with green-tinted midribs on the flowers – now known as Convallaria pseudomajalis. C. transcaucasica is recognised as a distinct species, subspecies, or variety of C. majalis by some authorities, but is not generally accepted as distinct. The species formerly called Convallaria japonica is now classified as Ophiopogon japonicus. ==Distribution==

Convallaria majalis is a native of Europe, where it largely avoids the Mediterranean margin, and is also absent from Ireland as a native species, though is naturalised there. ==Ecology==

Convallaria majalis is a plant of partial shade, and a mesophile type that prefers warm summers. It grows widely in both acidic soils and alkaline soils, liking soils that are silty or sandy, but also locally in wet fen soils, with preferably a plentiful amount of humus. It is a Euroasiatic and suboceanic species that occurs from sea level up to altitude in Great Britain, Convallaria majalis is used as a food plant by the larvae of some moth and butterfly (Lepidoptera) species including the grey chi. Adults and larvae of the leaf beetle Lilioceris merdigera are also able to tolerate the cardenolides and thus feed on the leaves. The fruit is sometimes removed by graniviorous rodents, consuming most of the seeds but only a small proportion of the fruit pulp. Their hoarding of both seeds and whole fruit has been observed. As some seeds inevitably escape predation, they also act as seed dispersers. ==Cultivars==

Convallaria majalis is widely grown in gardens for its scented flowers and ground-covering abilities in shady locations. It has gained the Royal Horticultural Society's Award of Garden Merit. In favourable conditions it can form large colonies. Various kinds and cultivars are grown, including those with double flowers, rose-coloured flowers, variegated foliage and ones that grow larger than the typical species. ==Chemistry==

Roughly 38 different cardiac glycosides (cardenolides) – which are highly toxic if consumed by humans or animals – occur in the plant, including: The scent of lily of the valley, specifically the ligand bourgeonal, was once thought to attract mammalian sperm. The 2003 discovery of this phenomenon prompted research into odour reception, but a 2012 study demonstrated instead that at high concentrations, bourgeonal imitated the role of progesterone in stimulating sperm to swim (chemotaxis), a process unrelated to odour reception. Toxicology All parts of the plant are potentially poisonous, including the red berries which may be attractive to children. If ingested, the plant can cause abdominal pain, nausea, vomiting, and irregular heartbeats. ==Uses==

Perfume In 1956, the French firm Dior produced a fragrance simulating lily-of-the-valley, which was Christian Dior's favourite flower. Diorissimo was designed by Edmond Roudnitska. Although it has since been reformulated, it is considered a classic. Because no natural aromatic extract can be produced from lily of the valley, its scent must be recreated synthetically; while Diorissimo originally achieved this with hydroxycitronellal, the European Chemicals Agency now considers it a skin sensitizer and its use has been restricted. Other perfumes imitating or based on the flower include Henri Robert's Muguet de Bois (1936), Penhaligon's Lily of the Valley (1976), Lily of the valley was featured in the bridal bouquet at the wedding of Prince William and Catherine Middleton. Lily of the valley was also the flower chosen by Princess Grace of Monaco to be featured in her bridal bouquet. At the beginning of the 20th century, it became tradition in France to sell lily of the valley on international Labour Day, 1 May (also called or Lily of the Valley Day) by labour organisations and private persons without paying sales tax (on that day only) as a symbol of spring. Lily of the valley is worn in Helston (Cornwall, UK) on Flora Day (8 May each year, see Furry Dance) representing the coming of "the May-o" and the summer. There is also a song sung in pubs around Cornwall (and on Flora Day in Cadgwith, near Helston) called "Lily of the Valley"; the song, strangely, came from the Jubilee Singers from Fisk University in Nashville, Tennessee. Folk medicine The plant has been used in folk medicine for centuries. There is a reference to "Lilly of the valley water" in Robert Louis Stevenson's 1886 novel Kidnapped, where it is said to be "good against the Gout", and that it "comforts the heart and strengthens the memory" and "restores speech to those that have the dumb palsey". There is no scientific evidence that lily of the valley has any effective medicinal uses for treating human diseases. ==Cultural symbolism==

The lily of the valley was the national flower of Yugoslavia, and it also became the national flower of Finland in 1967. In the "language of flowers", the lily of the valley signifies the return of happiness. Myths and religion The name "lily of the valley", like its correspondences in some other European languages, is apparently a reference to the Bible phrase "lily of the valleys" (sometimes also translated as "lily of the valley") in Song of Songs 2:1 (). European herbalists' use of the phrase to refer to a specific plant species seems to have appeared relatively late in the 16th or 15th century. The Neo-Latin term convallaria (coined by Carl Linnaeus) and, for example, the Swedish name '' derives from the corresponding phrase lilium convallium'' in the Vulgate. In culture The lily of the valley is represented both literally and symbolically in art, literature, music, and other media. The flower is the theme of a poem by Paul Laurence Dunbar, and Tchaikovsky wrote the poem "Lilies of the Valley" () in December 1878 while in Florence. In Anton Chekhov's 1898 short story "A Doctor's Visit", drops of convallaria are mentioned as medicine. Marc Chagall produced the painting Lillies-of-the-Valley in 1916. Lily of the valley was reputedly Queen Elizabeth II's favourite flower, and so it was the theme of the poem "Floral Tribute" by the Poet Laureate Simon Armitage, written in memory of the Queen and published in the week after her death. Previously, an eponymous song was written by English rock band Queen and released as part of the 1974 studio album Sheer Heart Attack. "Face Off", the finale of the fourth season of the television series Breaking Bad, features the use of berries from lily of the valley as poison because the symptoms are similar to ricin poisoning. In the third episode of Outlander, children are revealed to have been dying after confusing Lily of the Valley for garlic and eating it. ==Gallery==

File:Convallaria-oliv-r2.jpg|Convallaria close-up File:Convallaria majalis Kemi, Finland 02.06.2013.jpg|Convallarias in Kemi in early June File:Stamp of Moldova 429.gif|Moldovan stamp File:Pennä reverse Anu 1990.jpg|Finnish 10 penny coin with the Convallaria engraving File:1 of May, 1851.jpg|The First of May 1851 by Franz Xaver Winterhalter File:Lunner komm.svg|Lunner (Norway) municipal coat of arms ==See also==