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Joggins Formation

The Joggins Formation is a geologic formation in Nova Scotia. It preserves fossils dating back to the Westphalian stage or Moscovian stage of the Upper Carboniferous period or Pennsylvanian period, including Hylonomus, the earliest known reptile. In addition to fossils, the Joggins Formation was a valuable source of coal from the 17th century until the mid-20th century.

History
Early mining Prior to European colonization, the Joggins Formation and surrounding territory was part of the Miꞌkmaꞌki, the traditional homeland of the Mi'kmaq nation. French colonization city of the Bay of Fundy began in 1604. Acadian miners from Beaubassin were the first Europeans to mine the cliffs at Joggins, taking advantage of the deposits less than a decade after Jean-Baptiste-Louis Franquelin visited the site in 1686. Though Franquelin failed to make any mention of coal on his 1686 map, the document precisely detailed the geography of Chignecto Bay, and in a more detailed map published in 1702 he named the Joggins Cliffs "Ance au Charbon", or "Cove of Coal". The explorer Antoine de la Mothe Cadillac reported coal in Chignecto Bay in 1692. According to a travelogue written by Robert Hale in 1731, coal had been mined from the Fundy Seam - one of the thickest coal veins at Joggins - for at least thirty years at the time of his visit, and though there are only sporadic allusions to these pre-English operations in written records the government of Nova Scotia recognized Acadian brothers René and Bernard LeBlanc as being the first to discover coal at Joggins in 1701. Eastern Acadia was captured by Great Britain in the War of the Spanish Succession and became the colony of Nova Scotia; the territory was formally ceded by France in 1713's Peace of Utrecht. Though the British had likely been mining at Joggins since the occupation began in 1710, the first official record of a coal mine at Joggins was made in 1720, when Governor of Nova Scotia Richard Philipps complained that the trade of coal between Nova Scotia and New England was going unregulated. Captain Andrew Belcher was one of the first merchants to transport coal from the Acadian-run mines in Joggins to the city of Boston, which was in the grips of a coal shortage in the 1710s, and after Belcher's death in 1717 his son Jonathan Belcher continued to trade along this route. Henry Cope's mine Governor Richard Philipps approved the creation of a government-owned coal mine at Joggins in 1730, to test the feasibility of both mining the resource and loading onto ships at the site. The project was led by Major Henry Cope, who invested in the mine with his own money and enlisted the help of Boston merchants to help establish the mine at what is today known as "Coal Mine Point". Coal was first harvested from Cope's mine in April 1731, and the Nova Scotia Council agreed to fund the project on 24 June 1731. Rather than load ships on-site, Cope had the coal transported by small boats from Joggins to a wharf at the mouth of Gran'choggin (present-day Downing Cove), a creek north of the mine. The name "Gran'choggin" represents the earlier known use of the name that would give rise to "Joggins", and is likely either a Romanization of the Mi'kmaq word "chegoggins" (English: "great encampment") or a portmanteau of the French word "grand" (English: "large") and Mi'kmaq word "choggin" (English: "creek"). Tides in the Bay of Fundy made docking in the Gran'choggin wharf difficult. Robert Hale arrived at the mine on 25 June 1731, recording in his travelogue that it took nineteen days for his schooner, the Cupid, to travel from Boston to Joggins, but it wasn't until 26 June that the ship was able to dock at Gran'choggin. Hale observed that a single miner could produce many chaldrons of coal each day, and that the Cupid was only the third ship to be loaded with coal from Cope's mine. The Cupid departed Joggins on 30 June, loaded with sixty tons of coal to be sold in Boston. Henry Cope and his fellow developers were granted a land grant to develop a area around the mine on 21 June 1732. By the terms of the land grant, several names in the area were changed: the cliffs were renamed to "Adventurer's Clifts" and Gran'choggin to "New Castle Cove". The land grant also stipulated that Cope pay a tax of one shilling and sixpence for every chaldron mined, send coal to Annapolis Royal to support military fortifications there, and begin construction of a town that would be named "Williamstown". To further support the venture, Cope demanded his crew of miners - about a dozen local Acadians - pay him rent in exchange for living on his land and working in his mine. The men conspired with their Mi'kmaq neighbours, and a group of three Mi'kmaq men raided Cope's land in 1732. The mine, storehouses, and Stanwell Hall (the first house built by Cope at the site) were destroyed in the attack, setting back operations substantially. Cope was soon unable to make funding for the project stretch, and after he defaulted on paying wages to his workers the mine was abandoned in November 1732. With the support of Governor Richard Philipps and James Brydges, 1st Duke of Chandos, Cope attempted to restore the mine in January 1733. As a condition of his sponsors, Philipps was tasked with building blockhouses at the top of the cliffs to provide the site with military protection. Though at least some of the necessary fortifications were built, the project was again abandoned by Cope, ending all operations at the mine. British settlement British cartographers Edward Amherst and George Mitchell scouted the region in 1735, renaming the Adventurer's Clifts to "Grand Nyjagen". Lieutenant Governor of Nova Scotia Lawrence Armstrong was insistent on having the area colonized by British settlers as it would secure their dominance in the region, which was still largely populated by the Mi'kmaq and Acadians. On 30 August 1736 a land grant was approved for the area around the northernmost reaches of the Grand Nyjaden for the purposes of establishing a town named "Norwich" (present-day Minudie), but the community failed to be established and was dissolved on 21 April 1760. Also in 1736, Captain Thomas Durrel produced his own map of Chignecto Bay in which he referred to the cliffs as "Sea Coal Cliff" and New Castle Cove as "Grand Jogin". Acadian settlers continued to establish themselves in the region after the failed attempt to establish Norwich. In this period, the Acadians again began to mine for coal along the Grand Nyjagen cliffs and elsewhere, digging a line of coal pits from the coast to River Hebert, which came to be called the "Rivère des Mines des Hébert". A geographical description of Nova Scotia was published in The London Magazine in 1749. The article commented on the abundance of coal in the Bay of Fundy, referenced Henry Cope's failed mining project, and assured readers that "a better use will, doubtless, be made of this treasure, when Nova Scotia itself comes to be inhabited". took over nearly all Nova Scotian mining operations in 1827. In 1825, the Duke of York had been deeply in debt to Rundell, Bridge & Co. when the firm learned of the 1788 lease that would have granted him mineral rights to all of Nova Scotia. A mining engineer was sent by Rundell, Bridge & Co. to scout Nova Scotia for copper, and while little of it was found the engineer reported that the colony was rich in coal. At the firm's insistence, the 1788 lease was rewritten to include coal and approved by King George IV. According to the new terms, this granted the Duke mining rights to the following for the next sixty years: Once the lease was issued, the Duke subletted it to Rundell, Bridge & Co. in exchange for 25% of all profits they generated off the minerals it covered. Rundell, Bridge & Co. founded a new firm, the General Mining Company, which would manage their Nova Scotian properties from London. Taking advantage of the region's natural resources, particularly coal, was imperative to the growth of British North America: fuel was hard to come by, the anthracite mined in the Coal Region of Pennsylvania being difficult to use and bituminous coal from the Appalachian Mountains proving costly to transport. The Association focused initially on deposits in Pictou and Sydney, where they were able to buy leases on preexisting mines and take advantage of infrastructure that was already in place. After acquiring the rights to Cape Breton (which had to be negotiated for separately) the GMA had established a monopoly over all the mines in Nova Scotia, and the Nova Scotia House of Assembly discouraged competition by refusing to grant leases to new mines even if doing so would not violate the terms of the Duke of York's lease. Though Des Barres's land claim had expired after his death, the GMA did not immediately make an attempt to establish themselves at Joggins, and actively discouraged anyone else from mining the Joggins Formation. Despite their urgings, small groups of Cornish settlers continued mining exposed coal veins in the area. By 1836 an illegal mine had sprung up around the King's Vein, accessed through an adit dug into the cliff wall where an outcropping had once been apparent. Abraham Gesner returned to Nova Scotia in 1844 to petition the House of Assembly for the rights to mine at Joggins, as he felt the coal reserves had gone unused for too long and a shortage of firewood in Cumberland County provided ample opportunity for a new source of fuel. The colonial government, which had previously protected the General Mining Association from any threat of competition, had begun to change its stance on the GMA's monopoly: it was coming to light that the firm was offering discounts to American buyers but not Nova Scotians, and an 1842 report commissioned by Lieutenant Governor Lucius Cary suggested the GMA was poorly managing the Albion Mine in Pictou. The government decided to approve Gesner's request. To preserve their dominance in the colony, the GMA subleased a tract of land near the community of Joggins Mines (present-day Joggins) and opened the Joggins Mine in 1847. Joggins Mine was a shaft mine roughly deep, accessing the King's Vein from above and using a horse gin to raise the coal to the surface, where it was loaded onto a narrow-gauge railway and transported to a pier near Bell's Brook to be loaded onto ships. The GMA hired Joseph Smith, manager of their Pictou mine, to oversee its early operations. Much of the coal mined at Joggins Mine was sold to buyers in Saint John, New Brunswick. Joggins Mine cost £16,000 to build, and after the initial start up costs were covered the GMA invested minimally in the site. The General Mining Association lost many of its claims in 1858, but retained the right to mine from the Joggins Formation. By 1866, the Joggins Mine was producing an average of 8,478 chaldrons of coal per year, making it the least productive of any GMA mine in Nova Scotia. However, the Joggins Mine required only 9 horsepower of steam to operate normally and thus produced 943 chaldrons for each unit of horsepower expended, far more efficient than any other owned by the GMA. The lycopsid had fallen from the cliffs, and has previously been entombed in Coal 15. Lyell and Dawson also discovered fossilized millipedes in the lycopsid trunks, and fossils of the land snail Dendropupa. Among other things, the evidence of "reptile" (the distinction between reptiles and amphibians had yet to be made clear in 1852) fossils in strata of such age would disprove the argument made by proponents of catastrophism that Palaeozoic fishes had lost global dominance to the reptiles amid a worldwide catastrophe at the beginning of the Mesozoic era. Following the expedition, Charles Lyell took the fossils to be studied in Boston, where they were studied by Louis Agassiz and Jeffries Wyman at Harvard University. Agassiz first dismissed the remains as belonging to a coelacanth and Wyman believed they were those of a marine reptile related to Proteus anguinus, but after working the fossils out of the stone they'd been recovered in the scientists confirmed that the fossils represented bones from two reptiles: seven vertebrae and an ilium. At the time, only four tetrapod specimens had been recovered from coal formations around the world, meaning the eight bones recovered by Lyell and Dawson represented one-third of all Carboniferous tetrapod fossils. Lyell communicated this discovery to John W. Dawson in a letter on 6 November 1852, and publicly announced them in a series of lectures - the "Lowell lectures" - delivered at the Lowell Institute several weeks later. It was around this time that Lyell and Dawson became aware of William Logan's 1843 survey of the Joggins Formation, and on 16 December 1852 Lyell sent a request to Logan for a copy of his work to compare against their own. Logan insisted in a letter dated 10 January 1853 that he had already sent Lyell a copy years earlier when his report was first issued, illustrating the ambivalence the researchers had towards each other. Lyell and Dawson's discovery was relayed to the Geological Society in London on 19 January 1853, but unbeknownst to either of them a postscript had been added to their paper by Richard Owen, who had described and named the fossil as Dendrerpeton acadianum. In 1855, Dawson collected his and Lyell's research on Joggins into Acadian Geology, a text that drew serious interest from geologists across England and the United States. 1855 also marked the only trip that Othniel Charles Marsh made to Joggins. Marsh had only recently begun to study at Yale University and was likely drawn to Joggins by the abundance of fossils described by Charles Lyell and John W. Dawson. Marsh returned to Yale with two vertebrae, which he described in 1862 as belonging to the marine reptile Eosaurus acadianus. Six years later Dawson commented that Marsh's description of Eosaurus closely resembled that of Ichthyosaurus, which at the time had only been found in European limestone dated to the Early Jurassic such as the quarries of Lyme Regis, deposits which the Joggins Formation predates by more than 100 million years. Vertebrate palaeontologists Donald Baird, Alfred Romer, and Robert L. Carroll have suggested that Marsh's Eosaurus actually represents fossils of Ichthyosaurus which Marsh purchased off someone who claimed they had been found at Joggins and then passed the find off as his own discovery. Based on bivalve fossils recovered from the Joggins Formation, Joseph Frederick Whiteaves described Asthenodonta in 1893, but the genus was redescribed later that year by Thomas Chesmer Weston, who renamed it Archanodon. George Frederick Matthew, a pioneer in the field of ichnology also took an interest in Joggins and published his observations on tetrapod trackways recovered from the site in 1903. Walter A. Bell began researching the Joggins Formation in 1911, almost immediately after his graduation from Yale University. Bell was one of Canada's delegates at International Geological Congress's twelfth meeting in 1913, and accompanied the other delegates on a tour and dinner at the Joggins Cliffs that year. In 1914, Bell conducted the first detailed study of macrofloral fossils across Nova Scotia's Carboniferous formations, including the Mabou Group and Cumberland Group; as part of this study Bell assigned Divisions III, IV, and V (and part of Division II) to the "Joggins Formation", the first use of the name. The number of workers in Joggins steadily increased with production, but as Nova Scotia deindustrialised and other mines shut down the percentage of Cumberland County miners employed at Joggins increased from 21% in the 1880s to 25% in the 1900s and 43% in the 1910s. The Provincial Workmen's Association (PWA) established an early foothold in Joggins, first with Brunswick Lodge and later with Holdfast Lodge. Brunswick Lodge was the first to represent Joggins Mine workers. In 1884 the union attempted to negotiate with their employer at the time, prompting the manager - known in contemporary accounts as "Mr. B." - to remove his gloves and threaten to beat every worker. The company withdrew from negotiations and offered workers $10.00 each to abandon the union and come back to work. The strike went on, and when one miner attempted to retrieve his wages from Mr. B. the manager threw a piece of cast iron at him. The miner retaliated by throwing the iron back at Mr. B., striking him in the head but causing no severe injuries. Despite the clear belligerence between the company and the workers, strikers eventually began to cross the picket line and Brunswick Lodge dissolved that year. '' caricature of William Stevens Fielding, 1909 By 1895, Holdfast Lodge represented 20% of Joggins Mine's workers, the largest union in mainland Nova Scotia at the time. The politics of Holdfast Lodge were considered radical even by the standards of other Provincial Miner Workers chapters in Nova Scotia. 15 of the 19 PMW strikes that were held in the 1890s happened in Cumberland County, and Holdfast Lodge organized three in 1895 alone. Robert Drummond, Grand Secretary of the PMW, was particularly annoyed by their negotiations and tactics, their membership being largely uneducated and militant. Amid the threat of a wage reduction in the winter of 1895–96, Holdfast Lodge ordered all Joggins Mine workers to strike despite being ordered by Drummond not to, and in mid-March 1896 an estimated 200 members of Holdfast Lodge barricaded themselves inside the local PWA meeting hall to resist arrest, arming themselves with bricks, a number of assorted small arms, and 27 rifles. Nonetheless, Premier William Stevens Fielding engaged with Holdfast Lodge in order to build a loyal and powerful voter base. Fielding was leader of the Anti-Confederation Party and had been elected on a promise to remove Nova Scotia from Confederation, and when he failed to fulfill this promise he refocused his efforts on expanding the province's coal industry instead. One of Fielding's unconventional tactics involved working with the PWA to improve workers' access to education. The federal Liberal Party also campaigned hard for the PWA, as Cumberland had long been a Conservative Party stronghold: Cumberland had only ever been represented by Conservative Members of Parliament since Confederation and in nine elections had supported Charles Tupper, who was leading the Conservatives in 1896. Support from Holdfast Lodge managed to turn Cumberland in favour of the Liberals, and Hance James Logan was elected to the Parliament on 23 June 1896. The PWA succeeded in part because of their support for the community's funerary traditions, shutting down facilities like the Joggins Mine following a death and not returning to work until the deceased worker was put to rest. Such elaborate traditions had once been commonplace, but were mostly abandoned by the turn of the century until revived with the protection offered by unions. In 1898 the manager of Joggins Mine insisted the workers send a delegation to the funeral rather than shut down the mine for a day to attend; this request was ignored. In 1906, a funeral procession consisting of all the miners in Joggins marched from town to the Maccan River Bridge. The PWA came into conflict not just with mine managers, but also with the United Mine Workers, which had been founded in the United States in 1890. The PWA and the provincial United Mine Workers organization united in 1917 to form the Amalgamated Mine Workers of Nova Scotia, which was assimilated into the UMW in 1918. Child labour As was the case with many of Nova Scotia's mining operations, the coal mines at Joggins employed child labour when possible. Following the deaths of 70 miners and a single boy in the Drummond Mine explosion on 13 May 1873 in Westville, miners in Joggins adopted a rule of allowing children working in the mines to be the first to leave at the end of the day. While children were paid less than their adult colleagues, their income was indispensable to mining families which were typically impoverished. Class disparity became obvious with the advent of education as a status symbol, poor children often abandoning school early in life to work the mines. In August 1905 a number of child miners at Joggins left work on a recreational strike when they left work to play a game of baseball. Ideas surrounding childhood began to shift in the mid-19th century. The Mines Act of 1873 prohibited anyone under the age of 10 from working in any Nova Scotian mine, and in 1891 the minimum age of miners was raised to 12. In 1908 legislation banned the employment of any child miner who had not completed a grade seven education. Nova Scotia adopted the Free Schools Act in 1864 to provide public funding for schools which agreed to follow provincial standards of education, and in 1883 public school boards were given the right to fine parents of children aged 7–12 if their child did not attend at least 80 days of school per year. Nova Scotia finally enforced province-wide compulsory attendance for children aged 7–14 in 1921. In 1923 the Mines Act was amended to ban children from coal mines altogether, raising the school-leaving age to 16 and prohibiting anyone below that age from working in a mine. Don Reid and the Joggins Fossil Centre Donald R. "Don" Reid (1922–2016) was born on 29 May 1922 in Joggins, Nova Scotia. Reid's family was involved in the Joggins Mine, around which Joggins's economy was based, and as a teenager he was forced to leave school when his father sustained an injury in the mine and could no longer work. During this time Reid developed an interest in the Joggins Formation's abundance of fossils, collecting and studying them despite having no formal training as a palaeontologist. Don Reid was not the only person taking an interest in the site, and in 1972 a 1.6 km (1 mi) section of the Joggins Cliffs were protected under the Historical Objects Protection Act, which was repealed and replaced with the Special Places' Protection Act in 1980 and prohibited fossil collecting from Joggins or anywhere else in Nova Scotia without a permit. Though the risk of fossil poaching was not high in Joggins, the legislation ensured Reid remained one of only a handful of people authorized to research the cliffs. The Reid family opened a visitor centre on their property in 1989 to exhibit the vast collection, spurring the creation of the Joggins Fossil Centre at Coal Mine Point in Spring 1993. Reid and his assistant often gave personal tours of the Joggins Cliffs, and the Reid family claims that the Joggins Fossil Centre hosted visitors from more than 44 countries. Reid acquired the nickname "Keeper of the Cliffs" from the Joggins community during this period. Reid was instrumental in having UNESCO declare the Joggins Fossil Cliffs a World Heritage Site, which occurred on 7 July 2008. A number of institutions and societies honoured Don Reid for his contributions to palaeontology, particularly in the latter years of his life. In 2013, the Atlantic Geoscience Society granted Reid the Laing Ferguson Distinguished Service Award, and in 2016 Reid was inducted into the Order of Nova Scotia. In 2015, researchers announced that a new ichnofossil - a tetrapod trackway found in several formations of the Cumberland Group including the Joggins Formation - would be named in honour of Reid when the species was formally described. The trackway consists of footprints wide, and likely represent the Joggins Formation's largest tetrapod and top predator (possibly Baphetes). Recent history The Joggins Formation, which had been treated only as a member of the Cumberland Group since 1944, was formally reclassified as its own formation in 1991. The new classification used the measurements of William Logan's 1843 survey and contained Divisions IV and V, as well as the base of Division III. While the decision to define this section as a distinct formation has been upheld in later studies, the exact boundaries of the Joggins Formation have been debated. In 2005, the Joggins Formation was reformulated to consist of only Division IV and the limestone base of Division III. A bronze plaque was erected in Joggins in May 1992 to commemorate the 150-year anniversary of the Geological Survey of Canada's inception. The plaque is dedicated to the work of Sir William Logan. An international team of geologists remeasured the Joggins Formation in 2005, the first time such a procedure had been done since Charles Lyell and John W. Dawson's survey in 1853. To celebrate ten years as a UNESCO heritage site, the Joggins Fossil Institute hosted the first Joggins Research Symposium on 22 September 2018. A number of improvements were suggested to improve the site for research and educational purposes, including the construction of a storage facility for fossil lycopsids recovered from the cliffs which could bear tetrapod fossils, the development of machine learning software to better identify fossils at the site, and revision of the Special Places Protection Act. == Geology ==
Geology
The Joggins Formation overlies the Little River and Grand Anse Formations, and underlies the Polly Brook Formation. It is part of the Cumberland Group of geological formations, which extends from the early Namurian stage to the Westphalian stage of the Carboniferous period. The deposit represents a time when the region was dominated by a tropical rainforest, and consists of an enormous quantity of sedimentary rock. This material was deposited by rivers and flood water moving northwards, leaving behind sediment that subsided between two fault blocks: the Cobequid Mountains and Caledonia Mountain (present-day Caledonia Mountain, New Brunswick), both of which were active in the Carboniferous. the exact measurements of each division varied, as Logan used paced measurements to calculate distances. The divisions are numbered from top to bottom, with Division 1 containing the youngest strata and Division 8 containing the oldest. Division IV Division IV forms the basis of the Joggins Formation, with only a small amount of the overlying strata being considered part of the broader formation. Measured originally by William Edmond Logan in 1843, Division IV consists of a stretch of coal-bearing strata along the Joggins Cliffs. Division IV has an aggregate coal thickness of , contrasting with of limestone. Other divisions Division II contains predominately red sandstone and mudstone. Division II has not been considered part of the Joggins Formation since 1944. Division III contains fewer coal groups and more red sandstone than Division IV, which it overlays, and no limestone. In total, Division III includes 22 coal seams with an aggregate thickness of . The limestone base of Division III is considered part of the Joggins Formation, but the rest is not. Composed largely of grey sandstone, red sandstone, and red mudstone, Division V also bears some green shale and limestone. Division V contains no coal groups, and stretches from Lower Cove to South Reef. Division V was considered to be part of the Joggins Formation when the formation was defined in 1914 and 1991, but has been considered distinct since 2005. Coal Coal is abundant in the Joggins Formation and surrounding Joggins Cliffs, and was mined for centuries. There are 45 coal groups located in the formation. Coal Group 45 lies at the base of the Joggins Formation, and though the section assigned to this group stretches from start to finish, only of this actually represents basal coal. Two of the most heavily mined deposits of coal - the Fundy Seam and Dirty Seam - are part of the Joggins Formation. The Fundy Seam is thick, and is made of bituminous coal, while the Dirty Seam is thick and made of clastic-rich coal; they are present at above the formation's stratigraphic base. Other major seams in the Joggins Formation include the Joggins Seam (formerly the "King's Vein" and "Coal 7"), Queen's Seam ("Coal 8"), Forty Brine Seam ("Coal 20"), and Kimberly Seam ("Coal 14"). The majority of the Joggins Formation lies between Coal 34 and Coal 45, where coal is abundant. Coal harvested from Joggins was described as being "of inferior quality abounding with sulphur" in 1787. Fossil Lycopsids The Joggins Formation is of particular interest to geologists for its saturation with fossilized plants, one of the best-preserved coal forests known to science. Though often referred to as "trees", the large plants that made up the Joggins Formation's forest were lycopsid, which today only exist as club mosses. In the Carboniferous, lycopsids could grow as tall as with trunks nearly in diameter and came to resemble modern trees through convergent evolution. As trees do not constitute a taxonomic group, Carboniferous lycopsids are considered as plants as with any extant species of trees, despite being only distantly related. When the region was covered with marshland lycopods would put down rhizomes on the solid ground of newly made alluvial plains where young individuals faced little competition but were at risk of flooding. If the river banks burst before the forest floor had accumulated a significant amount of peat, the lycopsids would be swamped by sand carried overland in a crevasse splay, burying the trunk and killing the lycophyte as well as any other living thing in or around it. One such event of this kind buried the Fundy forest, a section of the Joggins Formation located in Cycle 6, around from the base of the formation. Once dead, the section of trunk not submerged in sand would rot and decay while the section buried underground had a chance at fossilization. The Joggins Formation is famous for these lycopsid trunks, as while they are valuable in their own right they also protected the carcasses of many animals which fossilized as well. It is unknown how creatures like Hylerpeton or Dendropupa came to be trapped in the trunks, but it has been suggested they were either using the lycopsids as dens, or they fell into the hollow trunks as the lycopsids were rotting and were buried when sediment collapsed into the interior. One trunk recovered from the Joggins Formation contained thirteen separate vertebrate specimens. A total of eleven tetrapod species have been discovered within fossil lycopsid trunks at Joggins. While greater quantities of sand would preserve taller trunks, the higher mass would crush the lycopsid rhizomes, which is why Stigmaria fossils are only found around shorter trunks. Scour hollows up to deep and long surround some well-preserved trunks. Though lycopsid trunks are among the best known plants from the Joggins Formation, the genus Calamites represents more fossils than almost any other organism preserved in the Joggins Cliffs. Resembling the modern-day horsetails they are related to, the stems Calamites plants typically grew to be in diameter and more than tall, though one specimen was found to be thick and nearly tall. Calamites grew mostly upright, spacing apart from each other and forming a thick forest floor. Usually, only the lower portion of Calamites plants was preserved, but their mostly-intact fossils suggest they were buried very quickly, likely by the same methods which preserved lycopsid trunks. Despite the comparative notoriety of Joggins's fossil lycopsids and Hylonomus, no specimens of Hylonomus have ever been recovered from inside a lycopsid trunk; this likely due to the incompleteness of the fossil record. Fossil lycopsids are most often discovered in intervals between the Coal 29 (the Fundy Seam) and Coal 35. ==Palaeobiology==
Palaeobiology
Dozens of tetrapod, invertebrate, and plant fossils have been recovered from the Joggins Formation. A diverse array of ichnofossils have also been found at Joggins, including vertebrate trackways, invertebrate trace fossils, tunnel structures, rhizoliths, and possibly wood borings. so this absence is not strong evidence against marine influence in at least some levels of the formation. Indeed, subtle evidence of low tides may occur in some levels of the Joggins deposits, and some levels were probably deposited in brackish water, as suggested by the fossils found therein. Marine influence is indicated by limestone beds in the lower part of the formation even contain brachiopods and fragments of echinoderms that were probably transported, but from a nearby source. The "Hebert beds" are located in Cycle 5, roughly from the base of the Joggins Formation. While the name of this area is an informal designation, the Hebert beds are of great value to palaeontologists for the number of fossils located at the site. It is believed that the Hebert beds once hosted deep watering holes, which sustained surrounding plants and animals during dry seasons and prolonged droughts. Fossils found at the Hebert beds include Archanodon and Dendropupa shells. ==Animals==
Animals
Amphibians Microconchida Arthropods Fish Molluscs Reptiles Embolomeres Synapsids Incertae sedis ==Plants==
Plants
Cycads Ferns Lycophytes Progymnosperms ==Protists==
Ichnogenera
Invertebrate ichnofossils Vertebrate ichnofossils == See also ==
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