The supergroup
Amorphea contains very diverse heterotrophic organisms, from the macroscopic
fungi and
animals to the unicellular
choanoflagellates and classical
amoebae. They frequently exhibit the ability to produce multinucleated cells, a trait considered ancestral to Amorphea. They are also capable of producing
pseudopodia, as does the closely related
CRuMs clade, forming the clade
Podiata. Amorphea is divided into two clades:
Amoebozoa, containing well-known amoebae and slime molds, and
Obazoa, containing animals, fungi, and their closest relatives. The relationship between these two clades was initially called 'Unikonta', due to a hypothesis where their common ancestor was a unikont, i.e., a eukaryote with just one flagellum. However, this hypothesis was refuted, as there are bikont amorpheans (e.g.,
Breviata anathema) and it likely is not an ancestral trait to have a single flagellum.
Obazoa The clade
Obazoa contains two small groups of flagellates, the
breviates and the
apusomonads, and the large clade
Opisthokonta, which contains animals, fungi, and their closest protist relatives. but this taxon is paraphyletic, as apusomonads are more closely related to opisthokonts. The taxonomy of apusomonads was expanded in a 2022 phylogenetic study that introduced many new genera. • Class
Breviatea , order
Breviatida , family
Breviatidae , genera
Breviata,
Lenisia,
Pygsuia,
Subulatomonas. • Class
Thecomonadea , order
Apusomonadida , family
Apusomonadidae . Genera not assigned to any subfamily:
Amastigomonas,
Multimonas,
Podomonas,
Catacumbia,
Cavaliersmithia. • Subfamily
Apusomonadinae , genera
Apusomonas (=
Rostromonas),
Manchomonas. • Subfamily
Thecamonadinae , genera
Chelonemonas,
Karpovia,
Singekia,
Thecamonas.
Opisthokonta Opisthokonts are divided into two branches:
Holozoa (animal-related), containing the
ichthyosporeans,
pluriformeans,
filastereans and
choanoflagellates; and
Nucletmycea or Holomycota (fungus-related), containing the
nucleariids and
opisthosporidians. These groups, minus opisthosporidians, were classified as different classes within the paraphyletic phylum Choanozoa in the system of Cavalier-Smith, now obsolete. Instead, Choanozoa is the name used for the clade containing choanoflagellates and animals. Opisthosporidians (aphelids, rozellids and microsporidians) are often studied as protists, but are also considered fungi by protistologists Both Holozoa and Nucletmycea have been proposed once as superkingdoms by a group of mycologists who classified nucleariids and fungi as kingdoms, but without any mention of holozoan kingdoms. •
Nucletmycea (=Holomycota ) • Order
Rotosphaerida (=Cristidiscoidida ), family
Nucleariidae (=Pompholyxophridae ), genera:
Fonticula,
Nuclearia,
Parvularia,
Pompholyxophrys,
Lithocolla,
Vampyrellidium,
Elaeorhanis,
Pinaciophora,
Rabdiophrys,
Rabdiaster,
Thomseniophora. • Kingdom
Fungi •
Holozoa . Genera
incertae sedis:
Tunicaraptor, • Class
Ichthyosporea (=Mesomycetozoea ) • Order
Dermocystida , family
Rhinosporidaceae , genera
Amphibiocystidium,
Amphibiothecum,
Chromosphaera,
Dermocystidium,
Rhinosporidium,
Sphaerothecum,
Valentines. • Order
Ichthyophonida (=Ichthyophonae ), family
Amoebidiidae , genera
Abeoforma,
Amoebidium,
Anurofeca,
Astreptonema,
Caullerya,
Creolimax,
Eccrinidus,
Enterobryus,
Enteropogon,
Ichthyophonus,
Palavascia,
Pseudoperkinsus,
Psorospermium,
Sphaeroforma. • Class
Pluriformea , order
Corallochytrida • Family
Corallochytriidae , genus
Corallochytrium . • Family
Syssomonadidae , •
Choanozoa • Class
Choanoflagellata (=Craspedomonadina , Craspedomonadaceae , Craspedophyceae , Craspédomonadophycidées , Craspedomonadophyceae , Choanomonadea , Choanoflagelliida , Choanoflagellatea , Choanomonada ) • Order
Craspedida , family
Salpingoecidae (=Codonosigidae ), genera
Astrosiga,
Aulomonas,
Barroeca,
Choanoeca,
Cladospongia,
Codonocladium,
Codonosigopsis,
Codosiga (=
Codonosiga),
Desmarella (=
Codonodesmus,
Kentrosiga),
Dicraspedella,
Diploeca,
Diplosiga,
Diplosigopsis,
Hartaetosiga,
Kentia,
Lagenoeca,
Microstomoeca,
Monosiga,
Mylnosiga,
Pachysoeca,
Proterospongia,
Salpingoeca,
Salpingorhiza,
Sphaeroeca,
Stagondoeca,
Stelexomonas,
Stylochromonas. • Order
Acanthoecida • Family
Acanthoecidae , genera
Acanthoeca,
Enibas,
Helgoeca,
Polyoeca,
Savillea. • Family
Stephanoecidae , genera
Acanthocorbis,
Amoenoscopa,
Apheloecion,
Bicosta,
Calliacantha,
Calotheca,
Campanoeca,
Campyloacantha,
Conion,
Cosmoeca,
Crinolina,
Crucispina,
Diaphanoeca,
Didymoeca,
Kakoeca,
Monocosta,
Nannoeca,
Parvicorbicula,
Thomsenella (="
Platypleura"),
Pleurasiga,
Polyfibula,
Saepicula,
Saroeca,
Spinoeca,
Spiraloecion,
Stephanacantha,
Stephanoeca,
Syndetophyllum. • Kingdom
Metazoa (=Animalia )
Amoebozoa The phylum
Amoebozoa contains around 2,400 species of primarily amoeboid protists. After the general 2019 revisions published by the International Society of Protistologists (ISOP), and testate amoebae. Phylum
Amoebozoa . Genera
incertae sedis:
Belonocystis,
Boveella,
Biomyxa,
Corallomyxa,
Gibbodiscus,
Hartmannia,
Malamoeba,
Malpighamoeba,
Oscillosignum,
Pseudothecamoeba,
Rhabdamoeba,
Schoutedamoeba,
Stereomyxa,
Subulamoeba,
Triaenamoeba,
Unda. • Order
Trichosida , family
Trichosphaeriidae (=Trichosidae ), genus
Trichosphaerium (=
Atrichosa). • Family
Microcoryciidae , genera
Amphizonella,
Diplochlamys,
Microcorycia*,
Parmulina*,
Penardochlamys*,
Zonomyxa*. • Order
Echinamoebida • Family
Echinamoebidae , genera
Echinamoeba,
Micriamoeba. • Family
Vermamoebidae , genus
Vermamoeba. •
Elardia • Family
Flabellulidae , genus
Flabellula (=
Paraflabellula). • Family
Gephyramoebidae , genus
Gephyramoeba.
Proplagiopyxis,
Protoplagiopyxis,
Protocucurbitella,
Pseudawerintzewia,
Pseudonebela,
Schoenbornia,
Sexangularia,
Suiadifflugia,
Trigonopyxis,
Wailesella, • Suborder
Phryganellina • Family
Phryganellidae , genus
Phryganella. • Family
Cryptodifflugiidae , genus
Cryptodifflugia. • Suborder
Organoconcha , family
Microchlamyiidae , genera
Microchlamys (=
Pseudochlamys),
Spumochlamys,
Pyxidicula. • Suborder
Glutinoconcha • Infraorder
Volnustoma , family
Heleoperidae , genera
Heleopera,
Metaheleopera. • Infraorder
Hyalospheniformes , family
Hyalospheniidae (=Nebelidae ), genera
Alabasta,
Alocodera,
Apodera,
Certesella,
Cornutheca,
Gibbocarina,
Hyalosphenia,
Longinebela,
Mrabella,
Nebela,
Padaungiella,
Planocarina,
Porosia,
Quadrulella. • Infraorder
Excentrostoma , family
Centropyxidae (incl.
Plagiopyxidae ), genera
Awerinzewia,
Bullinularia,
Centropyxis,
Frenopyxis,
Golemanskia,
Plagiopyxis. • Infraorder
Cylindrothecina , family
Cylindrifflugiidae , genus
Cylindrifflugia. • Infraorder
Longithecina • Family
Difflugiidae , genera
Difflugia,
Zivkovicia. • Family
Lesquereusiidae , genus
Lesquereusia. • Infraorder
Sphaerothecina • Family
Arcellidae , genera
Arcella,
Galeripora. • Family
Netzeliidae , genera
Cucurbitella,
Cyclopyxis,
Netzelia. • Class
Discosea • Subclass
Flabellinia • Superorder
Dermelia • Order
Mycamoebida , family
Mycamoebidae , genera
Mycamoeba,
Janelia,
Microglomus. • Order
Dermamoebida • Family
Dermamoebidae , genera
Dermamoeba,
Paradermamoeba,
Coronamoeba.
Thecamoeba,
Thecochaos. • Order
Dactylopodida , genera
Cunea,
Janickina,
Korotnevella,
Neoparamoeba,
Paramoeba,
Pseudoparamoeba,
Vexillifera. • Order
Vannellida , family
Vannellidae , genera
Clydonella,
Lingulamoeba,
Paravannella,
Pessonella,
Ripella,
Vannella. • Subclass
Centramoebia • Order
Acanthopodida (=Centramoebida ), genera
Acanthamoeba,
Balamuthia,
Luapeleamoeba,
Protacanthamoeba,
Dracoamoeba,
Vacuolamoeba. • Order
Himatismenida , genera
Cochliopodium,
Ovalopodium,
Parvamoeba. • Order
Pellitida , genera
Endostelium,
Gocevia,
Paragocevia,
Pellita. •
Evosea • Class
Cutosea , order
Squamocutida • Family
Squamamoebidae , genera
Armaparvus,
Squamamoeba. • Family
Sapocribridae , genus
Sapocribrum. • Family
Idionectidae , genus
Idionectes. • Class
Variosea . Genera not assigned to any lower clade:
Angulamoeba,
Arboramoeba,
Darbyshirella,
Dictyamoeba,
Ischnamoeba,
Heliamoeba,
Filamoeba,
Phalansterium. • Family
Flamellidae , genera
Flamella (=
Comandonia),
Telaepolella . • Order
Protosteliida (Protosteliales) , family
Protosteliidae (Protosteliaceae) , genus
Protostelium (incl.
Planoprotostelium). • Order
Fractovitellida • Family
Acramoebidae , genus
Acramoeba. • Family
Schizoplasmodiidae , genera
Ceratiomyxella,
Nematostelium,
Schizoplasmodium. • Family
Soliformoviidae . Genera:
Soliformovum ;
Grellamoeba . • Order
Cavosteliida (Cavosteliales) , family
Cavosteliidae (Cavosteliaceae) , genera
Cavostelium,
Schizoplasmodiopsis,
Tychosporium. • Order
Holomastigida (incl. Artodiscida ), family
Multiciliidae (incl. Artodiscidae ), genera
Artodiscus,
Multicilia. • Class
Archamoebea • Family
Tricholimacidae , genus
Tricholimax. • Order
Entamoebida , family
Entamoebidae , genera
Entamoeba,
Entamoebites. • Order
Pelobiontida • Suborder
Pelomyxina , family
Pelomyxidae , genera
Mastigella (=
Mastigamoeba),
Pelomyxa. • Suborder
Mastigamoebina • Family
Rhizomastixidae , genus
Rhizomastix. • Family
Mastigamoebidae , genera
Endamoeba*,
Endolimax,
Iodamoeba,
Mastigamoeba,
Mastigina*. •
Eumycetozoa (=Mycetozoa , Macromycetozoa ) • Class
Dictyostelea (Dictyosteliomycetes ). Genus unassigned to any order:
Synstelium. Genus
incertae sedis:
Coenonia. • Order
Acytosteliales • Family
Acytosteliaceae , genera
Acytostelium,
Rostrostelium,
Heterostelium. • Family
Cavenderiaceae , genus
Cavenderia. • Order
Dictyosteliales . Genus unassigned to any family:
Coremiostelium. • Family
Dictyosteliaceae , genera
Dictyostelium,
Polysphondylium. • Family
Raperosteliaceae , genera
Hagiwaraea,
Raperostelium,
Speleostelium,
Tieghemostelium. • Class
Ceratiomyxomycetes (Ceratiomyxea) , • Family
Lamprodermataceae (Lamprodermatidae) , genera
Lamproderma (incl.
Dichaeopsis,
Colloderma,
Elaeomyxa),
Collaria. • Family
Didymiaceae (Didymiidae) , genera
Diderma,
Didymium,
Lepidoderma,
Mucilago. • Family
Physaraceae (Physaridae) , genera
Craterium,
Leocarpus,
Fuligo,
Physarum (incl.
Badhamia),
Physarella,
Physarina,
Kelleromyxa. ==Archaeplastida==