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Caspian tiger

The Caspian tiger was a Panthera tigris tigris population native to eastern Turkey, northern Iran, Mesopotamia, the Caucasus around the Caspian Sea, Central Asia to northern Afghanistan and the Xinjiang region in Western China. It inhabited sparse forests and riverine corridors in this region until the 1970s. It was also present in Southern Russia until the Middle Ages. The Caspian tiger population was regarded as a distinct subspecies and assessed as extinct in 2003.

Taxonomy
Felis virgata was a scientific name used by Johann Karl Wilhelm Illiger in 1815 for the greyish tiger in the area surrounding the Caspian Sea. Tigris septentrionalis was the scientific name proposed by Konstantin Satunin in 1904 for a skull and mounted skins of tigers that were killed in the Lankaran Lowland in the 1860s. Felis tigris lecoqi and Felis tigris trabata were proposed by Ernst Schwarz in 1916 for tiger skins and skulls from Lop Nur and Ili River areas, respectively. In 1929, Reginald Innes Pocock subordinated the tiger to the genus Panthera. For several decades, the Caspian tiger was considered a distinct tiger subspecies. In 1999, the validity of several tiger subspecies was questioned. Most putative subspecies described in the 19th and 20th centuries were distinguished on basis of fur length and colouration, striping patterns and body size, hence characteristics that vary widely within populations. Morphologically, tigers from different regions vary little, and gene flow between populations in those regions is considered to have been possible during the Pleistocene. Therefore, it was proposed to recognize only two tiger subspecies as valid, namely P. t. tigris in mainland Asia, and P. t. sondaica in the Greater Sunda Islands and possibly in Sundaland. At the start of the 21st century, genetic studies were carried out using 20 tiger bone and tissue samples from museum collections and sequencing at least one segment of five mitochondrial genes. Results revealed a low amount of variability in the mitochondrial DNA in Caspian tigers; and that Caspian and Siberian tigers were remarkably similar, indicating that the Siberian tiger is the genetically closest living relative of the Caspian tiger. Phylogeographic analysis indicates that the common ancestor of Caspian and Siberian tigers colonized Central Asia via the GansuSilk Road region from eastern China less than 10,000 years ago, and subsequently traversed eastward to establish the Siberian tiger population in the Russian Far East. The Caspian and Siberian tigers were likely a single contiguous population until the early 19th century, but became isolated from another due to fragmentation and loss of habitat during the Industrial Revolution. In 2017, the Cat Specialist Group revised felid taxonomy and now recognizes the tiger populations in continental Asia as P. t. tigris. ==Characteristics==
Characteristics
Fur Photographs of skins of Caspian and Siberian tigers indicate that the main background colour of the Caspian tiger's fur varied and was generally brighter and more uniform than that of the Siberian tiger. The stripes were narrower, fuller and more closely set than those of tigers from Manchuria. The colour of its stripes was a mixture of brown or cinnamon shades. Pure black patterns were invariably found only on head, neck, the middle of the back and at the tip of the tail. Angular patterns at the base of the tail were less developed than those of Far Eastern populations. The contrast between the summer and winter coats was sharp, though not to the same extent as in Far Eastern populations. The winter coat was paler, with less distinct patterns. The summer coat had a similar density and hair length to that of the Bengal tiger, though its stripes were usually narrower, longer and closer set. It had the thickest fur amongst tigers, possibly due its occurrence in the temperate parts of Asia. Size The Caspian tiger ranked among the largest extant cat species along with the Siberian tiger. Maximum skull length of male Caspian tigers was , while that of females was . Its occiput was broader than of the Bengal tiger. ==Distribution and habitat==
Distribution and habitat
In the 19th century, tigers occurred in: • the Eastern Anatolia region, which is considered to have been the westernmost area where tigers occurred. The only confirmed record in Iraq dates to 1887 when a tiger was shot near Mosul, which is considered to have been a migrant from southeastern Turkey. • the extreme southeast of the Caucasus, such as in hilly and lowland forests of the Talysh Mountains, in the Lenkoran Lowlands, in the lowland forests of Göytəpə, Jalilabad, from where tigers moved into the eastern plains of the Trans-Caucasus up to the Don River basin; the Arasbaran and Zangezur Mountains of northwestern Persia. • Central Asia, such as in southwestern Turkmenia along the Atrek River and its tributaries, and the Sumbar and Chandyr Rivers; in the western and southwestern parts of Kopet-Dag; in the environs of Ashkabad in the northern foothills; in Afghanistan along the upper reaches of Hari-Rud at Herat, and along the jungles in the lower reaches of the river; around Tejen and Murgap and along the Kushka and Kashan rivers; in the Amu Darya basin as far the Aral Sea and along the entire coast of the Aral Sea; along the Syr-Darya to the Fergana Valley as far as Tashkent and the western spur of Talas Alatau; along the Chu and Ili Rivers; all along the southern shore of Lake Balkhash and northwards into the southern Altai Mountains, and to southeastern Transbaikal or Western Siberia in the east. In China, it occurred in the Tarim, Manasi River and Lop Nur basins. Wild boar was the numerically dominant ungulate in forested habitats, along watercourses, in reed beds and in thickets of the Caspian and Aral Seas. Where watercourses penetrated deep into desert areas, suitable wild pig and tiger habitat was often linear, only a few kilometers wide at most. Red and roe deer occurred in forests around the Black Sea to the western side and around the southern side of the Caspian Sea in a narrow belt of forest cover. Roe deer occurred in forested areas south of Lake Balkhash. Bactrian deer lived in the narrow belt of forest habitat on the southern border of the Aral Sea, and southward along the Syr-Darya and Amu Darya rivers. Local extinction The demise of the Caspian tiger began with the Russian colonisation of Turkestan during the late 19th century. Its extirpation was caused by several factors: • Tigers were killed by large parties of sportsmen and military personnel who also hunted tiger prey species such as the Bactrian deer, and middle asian Wild boar. This wild pig's range underwent a rapid decline between the middle of the 19th century and the 1930s due to overhunting, natural disasters, and diseases such as swine fever and foot-and-mouth disease, which caused large and rapid die-offs. By 1910, cotton plants were estimated to occupy nearly one-fifth of Turkestan's arable land, with about one half located in the Fergana Valley. Last sightings In Iraq, a tiger was killed near Mosul in 1887. In Turkey, a pair of tigers was allegedly killed in the area of Selçuk in 1943. Several tiger skins found in the early 1970s near Uludere indicated the presence of a tiger population in eastern Turkey. Questionnaire surveys conducted in this region revealed that one to eight tigers were killed each year until the mid-1980s, and that tigers likely had survived in the region until the early 1990s. Due to lack of interest, in addition to security and safety reasons, no further field surveys were carried out in the area. One of the last known tigers was shot in Golestan National Park in 1953. Another individual was sighted in Golestān Province in 1958. In Turkmenistan, the last known tiger was killed in January 1954 in the Sumbar River valley in the Kopet-Dag Range. It reportedly disappeared in the Manasi River basin in the Tian Shan Range west of Ürümqi in the 1960s. In Kazakhstan, the last Caspian tiger was recorded in 1948, in the environs of the Ili River, the last known stronghold in the region of Lake Balkhash. ==Behaviour and ecology==
Behaviour and ecology
attacking a tiger, from Roman Syria, which occupied parts of what is now Anatolia and Mesopotamia Caspian tigers possibly followed migratory ungulates from one pasture to another; wild boars and cervids probably formed their main prey base. In many regions of Central Asia, Bactrian deer and roe deer were important prey species, as well as Caspian red deer and goitered gazelle in Iran; Eurasian golden jackals, jungle cats, locusts, and other small mammals in the lower Amu Darya River area; Saiga antelopes, wild horses and Persian onagers in the Miankaleh Peninsula; Turkmenian kulans, Mongolian wild asses, and mountain sheep in the Zhana-Darya and around the Aral Sea; and Manchurian wapiti and moose in the area of Lake Baikal. They caught fish in flooded areas and irrigation channels. In winter, they frequently attacked dogs and livestock straying away from herds. They preferred drinking water from rivers, and drank from lakes in seasons when water was less brackish. In 1930, tigers moved up the Ili River after a decline of the wild boar population; in 1945, tigers followed a mass migration of wild boars along the shore of the Aral Sea from the Amu Darya delta into the lower reaches of the Syr Darya. Hunters who tracked tigers to Lake Balkhash observed a decrease in their numbers there, which coincided with the expansion of human activities and the migration of wild boar toward the Chu River. In the Karauzyak region of Uzbekistan, a witnessed battle ended with the deaths of both animals as a result of severe wounds. A similar outcome was recorded in the autumn of 1845 in the talysh forests. Two tigers in southwestern Tajikistan harbored 5–7 tapeworms (Taenia bubesei) in their small and large intestines. ==Conservation==
Conservation
In 1938, Tigrovaya Balka was the first protected area in Tajikistan established in the lower reaches of Vakhsh River between the Panj and Kofarnihon Rivers; it was apparently the last refuge of the Caspian tiger. A tiger was seen there in 1958. After 1947, tigers were legally protected in the Union of Soviet Socialist Republics. DNA from a tiger caught in northern Iran and housed at Moscow Zoo in the 20th century was used in the genetic test that established the Caspian tiger's close genetic relationship with the Siberian tiger. An individual was born in Brookfield Zoo Chicago on 7 May 1935 and was still living on 1 January 1948. Reintroduction project Stimulated by recent findings that the Siberian tiger is the closest relative of the Caspian tiger, discussions started as to whether the Siberian tiger could be appropriate for reintroduction into a safe place in Central Asia, where the Caspian tiger once roamed. The Amu Darya delta was suggested as a potential site for such a project. A feasibility study was initiated to investigate if the area is suitable, and if such an initiative would receive support from relevant decision makers. A viable tiger population of about 100 animals would require at least of large tracts of contiguous habitat, with rich prey populations. Such habitat is not currently available, and cannot be provided in the short term. The proposed region is therefore unsuitable for the reintroduction, at least at the current stage. In autumn 2023, two captive tigers from the Netherlands had arrived at the Ile-Balkhash Nature Reserve in Kazakhstan, and another three to four tigers are expected to be introduced from Russia. ==In culture==
In culture
In the Roman Empire, tigers and other large animals imported from Africa and Asia were used during gladiatorial games. In the Taurus Mountains, stone traps were used to capture leopards and tigers. In the Fables of Pilpay, the tiger is described as furious and avid to rule over wilderness. The babr (, tiger) features in Persian and Central Asian culture. The name "Babr Mazandaran" is sometimes given to a prominent wrestler. It possibly celebrates Odaenathus' victory over the Persians, the archer representing Odaenathus and the tigers the Persians; Odaenathus is about to be crowned with victory by the eagle flying above him. ==See also==
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